Chien C. Lee

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  • A well-camouflaged Borneo Rainbow Toad (Ansonia latidisca) scales a mossy tree trunk in the submontane rainforest of western Borneo. Although little is known about the life history of this rare species, it is an agile climber and is probably highly arboreal. Sarawak, Malaysia (Borneo).
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  • Without a single sighting in nearly 90 years, the Borneo Rainbow Toad (Ansonia latidisca) was considered possibly extinct and listed by Conservation International as one of the "10 Most Wanted Amphibians". The species was rediscovered in 2011 by a team of herpetologists from the University of Malaysia Sarawak, although it remains extremely rare and little is known of its ecology or behavior. This brings a glimmer of hope in a time of global mass amphibian extinctions. Sarawak, Malaysia (Borneo).
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  • Unseen since 1924, the Borneo Rainbow Toad (Ansonia latidisca) was previously listed as one of the "world's top 10 most wanted frogs" by Conservation International in their Global Search for Lost Frogs in 2010. It was rediscovered in Sarawak in 2011. Sarawak, Malaysia (Borneo).
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  • Borneo Frog-eating Snake (Stegonotus borneensis), juvenile. Sarawak, Malaysia (Borneo).
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  • Virgin rainforest in western Borneo. West Kalimantan, Indonesia.
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  • Setting sun and thundershowers over the rugged landscape of West Kalimantan, Borneo.
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  • A confusing array of small Microhylid frogs occur in Borneo and it is likely that many cryptic species such as this one (Nanohyla cf. perparva) await distinction. Sarawak, Malaysia (Borneo).
    cld1604051.jpg
  • Distinguished by its green/yellow eyes, Hose's Bush Frog (Philautus hosii) is endemic to Borneo and considered Near Threatened due to severe habitat loss. Adults are usually found perched on vegetation over streams in lowland rainforest. Sarawak, Malaysia (Borneo).
    cld1704736.jpg
  • A confusing array of small Microhylid frogs occur in Borneo and it is likely that many cryptic species such as this one (Nanohyla cf. perparva) await distinction. Sarawak, Malaysia (Borneo).
    cld1604026.jpg
  • The Gading Flying Frog (Leptomantis gadingensis), described as recently as 2005, is a small species known from only a few localities in western Borneo.  It breeds in lowland streams. Sarawak, Malaysia (Borneo).
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  • Wallace’s Flying Frog (Rhacophorus nigropalmatus) is one of the largest of all tree frogs in Borneo. It is capable of gliding down from the forest canopy by using its enlarged webbed feet as parachutes. Sarawak, Malaysia (Borneo).
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  • Orchid (Dendrobium sp.). Sarawak, Malaysia (Borneo).
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  • Alocasia puncakborneensis, a new Bornean endemic described in 2020. Sarawak, Malaysia (Borneo).
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  • Orchid (Nephelaphyllum pulchrum). Sarawak, Malaysia (Borneo).
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  • Long-tailed Grass Lizard (Takydromus sexlineatus). Sarawak, Malaysia (Borneo).
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  • The dainty White-eared Tree Frog (Feihyla kajau) is endemic to the rainforests of Borneo. Sarawak, Malaysia.
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  • Phaius subtrilobus. This large and showy terrestrial orchid is endemic to submontane forests in Borneo. Sarawak, Malaysia.
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  • A giant leaf katydid (Pseudophyllus hercules), one of the world's largest, rests in the rainforest understory. Active only at night, they use their superb camouflage to remain undetected by predators during the day. Sarawak, Malaysia (Borneo).
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  • Wallace's Flying Frog (Rhacophorus nigropalmatus), male. Sarawak, Malaysia (Borneo).
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  • Huntsman spider preying on camel cricket, with parasitic midge. Sarawak, Malaysia (Borneo).
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  • The White-eared Tree Frog (Feihyla kajau) lays its eggs in clusters on leaves overhanging small streams. When the tadpoles emerge they drop down into the water below. Sarawak, Malaysia (Borneo).
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  • Darkling beetle (Tetraphyllus sp.). Sarawak, Malaysia (Borneo).
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  • Orthostheneboea exotica, female (Previously Parastheneboea), a moss-mimicking stick insect endemic to Borneo. Sarawak, Malaysia.
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  • Leaf-mining beetle (Botryonopa sp.). Sarawak, Malaysia (Borneo).
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  • Translucent katydid (Lacipoda immunda), male concealing itself on the undersurface of a leaf. Sarawak, Malaysia (Borneo).
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  • Katydid (Tympanophyllum atroterminatum), male concealing itself by laying flat on the underside of a leaf of its food plant (Ficus lepicarpa) during the day. Sarawak, Malaysia (Borneo).
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  • Luminous Porecap (Favolaschia manipularis), releasing spores. Sarawak, Malaysia (Borneo).
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  • Touch-Me-Not Stick Insect (Epidares nolimetangere), male. Sarawak, Malaysia (Borneo).
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  • The bright colors on the hind wings of this tiger moth (Areas galactina, female) warn of its distasteful nature. Sarawak, Malaysia (Borneo).
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  • Tiger beetle (Therates sp.). Sarawak, Malaysia (Borneo).
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  • The tiny chlorophyll-free flowers of a Thismia (T. cf. filiformis), a mycoheterotrophic plant. Sarawak, Malaysia (Borneo).
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  • Cicadas (Huechys fusca), mating. Sarawak, Malaysia (Borneo).
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  • Edentistoma octosulcatum, a rare centipede that is believed to have a vegetarian diet. Sarawak, Malaysia (Borneo).
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  • Everett's Tree Toad (Rentapia everetti). Sarawak, Malaysia (Borneo).
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  • Lowland Dwarf Toad (Pelophryne signata), male vocalising. Sarawak, Malaysia (Borneo).
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  • An adult male Giant Golden Orbweaver (Nephila pilipes) is dwarfed by a gigantic female of the same species.  The male's tiny size allows him to approach the aggressive female unnoticed, or at least be considered nothing more than an insignificantly small prey item. Sarawak, Malaysia (Borneo).
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  • Long-legged Bush Frog (Philautus kakipanjang). Sarawak, Malaysia (Borneo).
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  • Darkling beetle (Cerogria sp.). Sarawak Malaysia (Borneo).
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  • Cat Gecko (Aeluroscalabotes felinus). These slow-moving and normally extremely placid reptiles only put on an aggressive display when their other defenses (camouflage and tail dropping) have failed them. Members of the ‘eyelid geckos’, they are the only representative of the family Eublepharidae in Borneo. Sarawak, Malaysia.
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  • Peter's Bent-toed Gecko (Cyrtodactylus consobrinus). Sarawak, Malaysia (Borneo).
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  • Under the cover of darkness, a cicada sheds its nymphal skin to emerge as a fully grown adult.  Sarawak, Malaysia (Borneo).
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  • Katydid (Lacipoda immunda), female. Sarawak, Malaysia (Borneo).
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  • Katydid (Hapalophyllum vrazi), female. Sarawak, Malaysia (Borneo).
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  • With highly toxic chemical defenses ranging from benzoquinones to cyanide compounds, millipedes are generally avoided by most predators. However, in nature no matter how well protected you are, everyone has their Achilles' heel. In this case we have the Ectrichodiinid bugs, better known as ' Millipede Assassins'. These specialized hunters feed exclusively on millipedes, stabbing them with a stout proboscis, immobilizing them with venom, and are undeterred by their prey's thick armor and toxins. It is suspected that some species may actually be able to sequester the chemicals of the millipede to incorporate into their own defenses, but this remains to be studied. Sarawak, Malaysia (Borneo).
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  • Stick insect (Dinophasma saginatum), mating. Sarawak, Malaysia (Borneo).
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  • Wallace's Flying Frog (Rhacophorus nigropalmatus), male. Sarawak, Malaysia (Borneo).
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  • Dragon-headed Katydid (Lesina blanchardi), juvenile. Sarawak, Malaysia (Borneo).
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  • Mating pair of stick insects (Staelonchodes sodalis) showing extreme dimorphism between male and female of the same species. Sarawak, Malaysia (Borneo).
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  • A Sunda Pangolin (Manis javanica) forages at night for ant and termite nests, using its keen sense of smell to locate a nest. Sarawak, Malaysia (Borneo).
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  • Darkling beetle (Strongylium sp.). Sarawak, Malaysia (Borneo).
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  • Leaf-mining beetle (Anisodera sp.), mating. Sarawak, Malaysia (Borneo).
    cld1604654.jpg
  • The rigid armour-like scales of the Pangolin (Manis javanica) afford it a high degree of protection from predators.  The scales are considered as valuable in Chinese medicine, and the animal is frequently hunted for this reason. Sarawak, Malaysia (Borneo).
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  • Giant ladybird beetle (Synonycha grandis). Sarawak, Malaysia (Borneo).
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  • Lanjak Bush Frog (Philautus refugii). Sarawak, Malaysia (Borneo).
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  • Kuhl's Gliding Gecko (Gekko kuhli). By means of its large webbed feet and lateral skin flaps, this arboreal gecko is capable of gliding or parachuting between trees. Sarawak, Malaysia (Borneo).
    cld1701921.jpg
  • Kuhl's Gliding Gecko (Gekko kuhli). Equipped with webbed feet and parasail-like flaps of skin all along its body, this tree-dwelling gecko is able to ‘parachute’ when leaping through the air to escape predators. This not only softens its landing but also enables it to glide a considerable distance. Sarawak, Malaysia (Borneo).
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  • Pallid Monkey Moth (Sphingognatha asclepiades). Sarawak, Malaysia (Borneo).
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  • Within the microcosm of a mossy tree stump, a tiny hidden predator lies patiently in wait. Still a juvenile, this praying mantis (Haania sp.) will eventually mature and shed most of its moss-like camouflage in favor of fully developed wings. Sarawak, Malaysia (Borneo).
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  • Dwarf Toad (Pelophryne signata). Sarawak, Malaysia (Borneo).
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  • Wallace's Flying Frog (Rhacophorus nigropalmatus). Sarawak, Malaysia (Borneo).
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  • Red-legged Frog (Leptomantis rufipes). Sarawak, Malaysia (Borneo).
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  • A recently-germinated Dipterocarp tree seedling (Shorea sp.) reaches for the sunlight on the Bornean rainforest floor.
    cld10071556.jpg
  • Without a single sighting in nearly 90 years, the Borneo Rainbow Toad (Ansonia latidisca) was considered possibly extinct and listed by Conservation International as one of the "10 Most Wanted Amphibians". The species was rediscovered in 2011 by a team of herpetologists from the University of Malaysia Sarawak, although it remains extremely rare and little is known of its ecology or behavior. This brings a glimmer of hope in a time of global mass amphibian extinctions. Sarawak, Malaysia (Borneo).
    cld1701360.jpg
  • Perhaps one of the most beautiful of all Borneo's frogs: the Borneo Flying Frog (Rhacophorus borneensis). These gliding amphibians spend most of their life in the tree canopy, only rarely descending to ground level. Sarawak, Malaysia (Borneo).
    cld1724123.jpg
  • Perhaps one of the most beautiful of all Borneo's frogs: the Borneo Flying Frog (Rhacophorus borneensis). These gliding amphibians spend most of their life in the tree canopy, only rarely descending to ground level. Sarawak, Malaysia (Borneo).
    cld1724109.jpg
  • Like other small animals of the forest understory, this Borneo Forest Dragon (Gonocephalus bornensis) must juggle his attention between searching for prey (insects and spiders) while keeping a watchful eye out for larger predators. Although this species is unable to glide like the closely-related Dracos, it is nevertheless an agile climber and can move swiftly up and down trees. The impressive mane on the back of this male signifies his maturity - we will readily defend his territory against rival males. Danum Valley, Sabah, Malaysia (Borneo).
    cld1928123.jpg
  • Borneo Necklaced Partridge (Tropicoperdix graydoni), considered by some to be a subspecies of Chestnut-necklaced Partridge (Tropicoperdix charltonii) from mainland Southeast Asia. Sabah, Malaysia (Borneo).
    cld1824750.jpg
  • Borneo Anglehead Lizard (Gonocephalus bornensis). Sarawak, Malaysia (Borneo).
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  • Spending the majority of its time underground in burrows, the Borneo Narrowmouth Toad (Gastrophrynoides borneensis) is a rarely seen species and little is known of its life history. Despite its common name and unusual appearance, this frog is a member of the Microhylidae and not a true toad. Sarawak, Malaysia (Borneo).
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  • Borneo Horned Frog (Pelobatrachus nasutus), female, with Matang Narrow-mouthed Frog (Microhyla nepenthicola) - comparison between one of the largest and smallest frogs in Borneo. Sarawak, Malaysia (Borneo).
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  • Borneo Horned Frog (Pelobatrachus nasutus). Sabah, Malaysia (Borneo).
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  • Borneo Short-tailed Python (Python breitensteini). Sarawak, Malaysia (Borneo).
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  • Borneo Opposite-fingered Tree Frog (Feihyla inexpectata), male. Sabah, Malaysia (Borneo).
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  • Borneo Pygmy Elephant (Elephas maximus borneensis). Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Borneo Forest Dragon (Gonocephalus bornensis). Sabah, Malaysia (Borneo).
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  • Dense mist often cloaks Borneo's rainforest at dawn, this being the water vapor rising from the transpiration of the forest itself. Sabah, Malaysia (Borneo).
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  • A twisted liana winds its way into the canopy among the virgin rainforest of Tawau Hills National Park in northern Borneo. Home to some of the tallest tropical trees in the world, the forests in these hills is inhabited by a rich diversity of wildlife. Sabah, Malaysia (Borneo).
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  • Heavy rain falls over a small river amid lowland rainforest. Many inland regions of Borneo such as this receive as much as 4 meters of rainfall per year. Sarawak, Malaysia (Borneo).
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  • Although the canopy of the Borneo rainforest is bathed in the heat of the equatorial sun, as little as 5 percent of sunlight reaches the forest floor. Sabah, Malaysia (Borneo).
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  • Rising mist spills over the forested plateau of Maliau Basin, a large conservation area in the remote interior of northern Borneo. Sabah, Malaysia (Borneo).
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  • View of canopy of lowland mixed dipterocarp forest during a mass flowering event. Sabah, Malaysia (Borneo).
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  • Reds, oranges, yellows, and other bright colors among the rainforest canopy are a sign of a mass flowering event. Borneo's rainforests undergo a cyclic mass flowering every 4-6 years (usually triggered by a dry spell), during which the majority of the trees flower and fruit simultaneously. This periodic event is a fundamental factor in the ecology of nearly all rainforest organisms in Borneo.
    cld1909709.jpg
  • After several months of feeding on Aristolochia leaves, the mature caterpillar of this large birdwing butterfly (Troides andromache) prepares for its final moult into the pupal stage when metamorphosis will transform it into an adult. Sabah, Malaysia (Borneo).
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  • Sunrise and mist over virgin rainforest in northern Borneo. Danum Valley Conservation Area, Sabah, Malaysia.
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  • An aerial perspective of northern Borneo's lowland rainforest reveals the heterogeneity and incredible diversity of trees that comprise this ecosystem. Various flushes of reds, oranges, and yellows among the canopy as seen here are the first indications of a periodic mass flowering event during which as many as 80% of the trees may bloom simultaneously before setting fruit. Sabah, Malaysia (Borneo).
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  • Mossy cloud forest occurs at higher elevations on nearly all of Borneo's mountains. These habitats are characterized by high rainfall, frequent fog, cooler temperatures, and small trees which are covered with moss and a myriad of epiphytic plants. Sarawak, Malaysia (Borneo).
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  • Batang Ai National Park. Sarawak, Malaysia (Borneo).
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  • Sunlight filters through lowland rainforest into the entrance of a limestone cave in Mulu National Park. Sarawak, Malaysia (Borneo).
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  • Oil Palm (Elaeis guineensis) plantation showing small riparian buffer zones along rivers. Northern Borneo.
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  • Scat of Mountain Treeshrew (Tupaia montana) on the peristome of Nepenthes rajah. This will be washed into the pitcher with rain where it becomes a vital source of nutrients for the plant. Sabah, Malaysia (Borneo).
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  • Sunrise and mist over virgin rainforest in northern Borneo. Danum Valley Conservation Area, Sabah, Malaysia.
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  • Sometimes referred to as 'durian kura-kura' (tortoise durian), Durio testudinarum is one of the rarest of edible durian species. It flowers near the base of the tree and produces clusters of small fruits that bear a musky odor. Like other durians, the blooms of D. testudinarum are open at night, though the pollinator of this species remains unknown. Sarawak, Malaysia (Borneo).
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  • This pond amid lowland forest is a prime breeding habitat for many species of amphibians. Sarawak, Malaysia (Borneo).
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  • Weathering from rain and wind has given rise to these razor-sharp limestone pinnacles, which can be found on the upper slopes of Mount Api in Mulu National Park. Sarawak, Malaysia (Borneo).
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  • A tannin-colored stream meanders through freshwater swamp forest in Sarawak. Trees with stilt roots, pneumatophores, and buttresses, are more abundant in this waterlogged habitat, which is frequently flooded by rains. Sarawak, Malaysia (Borneo).
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  • An aerial view reveals the thick mist rising from the canopy after heavy rains on virgin rainforest in northern Borneo. These forests often receive over 4000ml of rain annually. Sabah, Malaysia.
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  • Large granite boulders decorate the coastline at Tanjung Datu National Park at the western tip of Borneo. Sarawak, Malaysia.
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  • Sunrise and mist over virgin rainforest in northern Borneo. Danum Valley Conservation Area, Sabah, Malaysia.
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  • Large stretches of virgin rainforest still remain in the remote interior of eastern Borneo. East Kalimantan, Indonesia.
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  • Unidentified mushrooms. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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