Chien C. Lee

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  • Without a single sighting in nearly 90 years, the Borneo Rainbow Toad (Ansonia latidisca) was considered possibly extinct and listed by Conservation International as one of the "10 Most Wanted Amphibians". The species was rediscovered in 2011 by a team of herpetologists from the University of Malaysia Sarawak, although it remains extremely rare and little is known of its ecology or behavior. This brings a glimmer of hope in a time of global mass amphibian extinctions. Sarawak, Malaysia (Borneo).
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  • A well-camouflaged Borneo Rainbow Toad (Ansonia latidisca) scales a mossy tree trunk in the submontane rainforest of western Borneo. Although little is known about the life history of this rare species, it is an agile climber and is probably highly arboreal. Sarawak, Malaysia (Borneo).
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  • Without a single sighting in nearly 90 years, the Borneo Rainbow Toad (Ansonia latidisca) was considered possibly extinct and listed by Conservation International as one of the "10 Most Wanted Amphibians". The species was rediscovered in 2011 by a team of herpetologists from the University of Malaysia Sarawak, although it remains extremely rare and little is known of its ecology or behavior. This brings a glimmer of hope in a time of global mass amphibian extinctions. Sarawak, Malaysia (Borneo).
    cld1604545.jpg
  • Unseen since 1924, the Borneo Rainbow Toad (Ansonia latidisca) was previously listed as one of the "world's top 10 most wanted frogs" by Conservation International in their Global Search for Lost Frogs in 2010. It was rediscovered in Sarawak in 2011. Sarawak, Malaysia (Borneo).
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  • Borneo Frog-eating Snake (Stegonotus borneensis), juvenile. Sarawak, Malaysia (Borneo).
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  • Virgin rainforest in western Borneo. West Kalimantan, Indonesia.
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  • Setting sun and thundershowers over the rugged landscape of West Kalimantan, Borneo.
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  • Horsfield's Flying Gecko (Gekko horsfieldii). Sarawak, Malaysia (Borneo).
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  • Dusky Mock Viper (Psammodynastes pulverulentus). Sarawak, Malaysia (Borneo).
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  • A confusing array of small Microhylid frogs occur in Borneo and it is likely that many cryptic species such as this one (Nanohyla cf. perparva) await distinction. Sarawak, Malaysia (Borneo).
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  • Distinguished by its green/yellow eyes, Hose's Bush Frog (Philautus hosii) is endemic to Borneo and considered Near Threatened due to severe habitat loss. Adults are usually found perched on vegetation over streams in lowland rainforest. Sarawak, Malaysia (Borneo).
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  • Wallace’s Flying Frog (Rhacophorus nigropalmatus) is one of the largest of all tree frogs in Borneo. It is capable of gliding down from the forest canopy by using its enlarged webbed feet as parachutes. Sarawak, Malaysia (Borneo).
    cld1702577.jpg
  • A confusing array of small Microhylid frogs occur in Borneo and it is likely that many cryptic species such as this one (Nanohyla cf. perparva) await distinction. Sarawak, Malaysia (Borneo).
    cld1604026.jpg
  • The Gading Flying Frog (Leptomantis gadingensis), described as recently as 2005, is a small species known from only a few localities in western Borneo.  It breeds in lowland streams. Sarawak, Malaysia (Borneo).
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  • Orchid (Dendrobium sp.). Sarawak, Malaysia (Borneo).
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  • Alocasia puncakborneensis, a new Bornean endemic described in 2020. Sarawak, Malaysia (Borneo).
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  • Long-tailed Grass Lizard (Takydromus sexlineatus). Sarawak, Malaysia (Borneo).
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  • The dainty White-eared Tree Frog (Feihyla kajau) is endemic to the rainforests of Borneo. Sarawak, Malaysia.
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  • Translucent katydid (Lacipoda immunda), male concealing itself on the undersurface of a leaf. Sarawak, Malaysia (Borneo).
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  • Luminous Porecap (Favolaschia manipularis), releasing spores. Sarawak, Malaysia (Borneo).
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  • Huntsman spider preying on camel cricket, with parasitic midge. Sarawak, Malaysia (Borneo).
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  • Edentistoma octosulcatum, a rare centipede that is believed to have a vegetarian diet. Sarawak, Malaysia (Borneo).
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  • Orchid (Nephelaphyllum pulchrum). Sarawak, Malaysia (Borneo).
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  • The White-eared Tree Frog (Feihyla kajau) lays its eggs in clusters on leaves overhanging small streams. When the tadpoles emerge they drop down into the water below. Sarawak, Malaysia (Borneo).
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  • Dwarf Toad (Pelophryne signata). Sarawak, Malaysia (Borneo).
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  • Peter's Bent-toed Gecko (Cyrtodactylus consobrinus). Sarawak, Malaysia (Borneo).
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  • Phaius subtrilobus. This large and showy terrestrial orchid is endemic to submontane forests in Borneo. Sarawak, Malaysia.
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  • A giant leaf katydid (Pseudophyllus hercules), one of the world's largest, rests in the rainforest understory. Active only at night, they use their superb camouflage to remain undetected by predators during the day. Sarawak, Malaysia (Borneo).
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  • Katydid (Tympanophyllum atroterminatum), male concealing itself by laying flat on the underside of a leaf of its food plant (Ficus lepicarpa) during the day. Sarawak, Malaysia (Borneo).
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  • Katydid (Hapalophyllum vrazi), female. Sarawak, Malaysia (Borneo).
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  • Touch-Me-Not Stick Insect (Epidares nolimetangere), male. Sarawak, Malaysia (Borneo).
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  • The bright colors on the hind wings of this tiger moth (Areas galactina, female) warn of its distasteful nature. Sarawak, Malaysia (Borneo).
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  • Tiger beetle (Therates sp.). Sarawak, Malaysia (Borneo).
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  • Wallace's Flying Frog (Rhacophorus nigropalmatus), male. Sarawak, Malaysia (Borneo).
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  • Cicadas (Huechys fusca), mating. Sarawak, Malaysia (Borneo).
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  • Everett's Tree Toad (Rentapia everetti). Sarawak, Malaysia (Borneo).
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  • Dragon-headed Katydid (Lesina blanchardi), juvenile. Sarawak, Malaysia (Borneo).
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  • Lowland Dwarf Toad (Pelophryne signata), male vocalising. Sarawak, Malaysia (Borneo).
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  • A Sunda Pangolin (Manis javanica) forages at night for ant and termite nests, using its keen sense of smell to locate a nest. Sarawak, Malaysia (Borneo).
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  • An adult male Giant Golden Orbweaver (Nephila pilipes) is dwarfed by a gigantic female of the same species.  The male's tiny size allows him to approach the aggressive female unnoticed, or at least be considered nothing more than an insignificantly small prey item. Sarawak, Malaysia (Borneo).
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  • Long-legged Bush Frog (Philautus kakipanjang). Sarawak, Malaysia (Borneo).
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  • Kuhl's Gliding Gecko (Gekko kuhli). By means of its large webbed feet and lateral skin flaps, this arboreal gecko is capable of gliding or parachuting between trees. Sarawak, Malaysia (Borneo).
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  • Darkling beetle (Strongylium sp.). Sarawak, Malaysia (Borneo).
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  • Darkling beetle (Cerogria sp.). Sarawak Malaysia (Borneo).
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  • Kuhl's Gliding Gecko (Gekko kuhli). Equipped with webbed feet and parasail-like flaps of skin all along its body, this tree-dwelling gecko is able to ‘parachute’ when leaping through the air to escape predators. This not only softens its landing but also enables it to glide a considerable distance. Sarawak, Malaysia (Borneo).
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  • Darkling beetle (Tetraphyllus sp.). Sarawak, Malaysia (Borneo).
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  • Wallace's Flying Frog (Rhacophorus nigropalmatus). Sarawak, Malaysia (Borneo).
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  • Orthostheneboea exotica, female (Previously Parastheneboea), a moss-mimicking stick insect endemic to Borneo. Sarawak, Malaysia.
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  • Leaf-mining beetle (Botryonopa sp.). Sarawak, Malaysia (Borneo).
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  • Under the cover of darkness, a cicada sheds its nymphal skin to emerge as a fully grown adult.  Sarawak, Malaysia (Borneo).
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  • Huntsman spider (Heteropoda sp.) cannibalizing on another huntsman spider. Sarawak, Malaysia (Borneo).
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  • With highly toxic chemical defenses ranging from benzoquinones to cyanide compounds, millipedes are generally avoided by most predators. However, in nature no matter how well protected you are, everyone has their Achilles' heel. In this case we have the Ectrichodiinid bugs, better known as ' Millipede Assassins'. These specialized hunters feed exclusively on millipedes, stabbing them with a stout proboscis, immobilizing them with venom, and are undeterred by their prey's thick armor and toxins. It is suspected that some species may actually be able to sequester the chemicals of the millipede to incorporate into their own defenses, but this remains to be studied. Sarawak, Malaysia (Borneo).
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  • The tiny chlorophyll-free flowers of a Thismia (T. cf. filiformis), a mycoheterotrophic plant. Sarawak, Malaysia (Borneo).
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  • Stick insect (Dinophasma saginatum), mating. Sarawak, Malaysia (Borneo).
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  • Lanjak Bush Frog (Philautus refugii). Sarawak, Malaysia (Borneo).
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  • Wallace's Flying Frog (Rhacophorus nigropalmatus), male. Sarawak, Malaysia (Borneo).
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  • Mating pair of stick insects (Staelonchodes amaurops) showing extreme dimorphism between male and female of the same species. Sarawak, Malaysia (Borneo).
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  • Red-legged Frog (Leptomantis rufipes). Sarawak, Malaysia (Borneo).
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  • Cat Gecko (Aeluroscalabotes felinus). These slow-moving and normally extremely placid reptiles only put on an aggressive display when their other defenses (camouflage and tail dropping) have failed them. Members of the ‘eyelid geckos’, they are the only representative of the family Eublepharidae in Borneo. Sarawak, Malaysia.
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  • Leaf-mining beetle (Anisodera sp.), mating. Sarawak, Malaysia (Borneo).
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  • Pallid Monkey Moth (Sphingognatha asclepiades). Sarawak, Malaysia (Borneo).
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  • Within the microcosm of a mossy tree stump, a tiny hidden predator lies patiently in wait. Still a juvenile, this praying mantis (Haania sp.) will eventually mature and shed most of its moss-like camouflage in favor of fully developed wings. Sarawak, Malaysia (Borneo).
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  • The rigid armour-like scales of the Pangolin (Manis javanica) afford it a high degree of protection from predators.  The scales are considered as valuable in Chinese medicine, and the animal is frequently hunted for this reason. Sarawak, Malaysia (Borneo).
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  • Giant ladybird beetle (Synonycha grandis). Sarawak, Malaysia (Borneo).
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  • Katydid (Lacipoda immunda), female. Sarawak, Malaysia (Borneo).
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  • Feather-legged Assassin Bug (Ptilocerus venosus). Members of this subfamily (Holoptilinae) are specialist predators on ants, some species even having a glandular trichome on their abdomens which serves to attract prey. Sarawak, Malaysia (Borneo).
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  • A recently-germinated Dipterocarp tree seedling (Shorea sp.) reaches for the sunlight on the Bornean rainforest floor.
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  • Perhaps one of the most beautiful of all Borneo's frogs: the Borneo Flying Frog (Rhacophorus borneensis). These gliding amphibians spend most of their life in the tree canopy, only rarely descending to ground level. Sarawak, Malaysia (Borneo).
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  • Perhaps one of the most beautiful of all Borneo's frogs: the Borneo Flying Frog (Rhacophorus borneensis). These gliding amphibians spend most of their life in the tree canopy, only rarely descending to ground level. Sarawak, Malaysia (Borneo).
    cld1724123.jpg
  • Borneo Anglehead Lizard (Gonocephalus bornensis). Sarawak, Malaysia (Borneo).
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  • Like other small animals of the forest understory, this Borneo Forest Dragon (Gonocephalus bornensis) must juggle his attention between searching for prey (insects and spiders) while keeping a watchful eye out for larger predators. Although this species is unable to glide like the closely-related Dracos, it is nevertheless an agile climber and can move swiftly up and down trees. The impressive mane on the back of this male signifies his maturity - we will readily defend his territory against rival males. Sabah, Malaysia (Borneo).
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  • Spending the majority of its time underground in burrows, the Borneo Narrowmouth Toad (Gastrophrynoides borneensis) is a rarely seen species and little is known of its life history. Despite its common name and unusual appearance, this frog is a member of the Microhylidae and not a true toad. Sarawak, Malaysia (Borneo).
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  • Borneo Horned Frog (Pelobatrachus nasutus). Sabah, Malaysia (Borneo).
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  • Borneo Short-tailed Python (Python breitensteini). Sarawak, Malaysia (Borneo).
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  • Borneo Horned Frog (Pelobatrachus nasutus), female, with Matang Narrow-mouthed Frog (Microhyla nepenthicola) - comparison between one of the largest and smallest frogs in Borneo. Sarawak, Malaysia (Borneo).
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  • Borneo Pygmy Elephant (Elephas maximus borneensis). Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Borneo Forest Dragon (Gonocephalus bornensis). Sabah, Malaysia (Borneo).
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  • Borneo Opposite-fingered Tree Frog (Feihyla inexpectata), male. Sabah, Malaysia (Borneo).
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  • Heavy rain falls over a small river amid lowland rainforest. Many inland regions of Borneo such as this receive as much as 4 meters of rainfall per year. Sarawak, Malaysia (Borneo).
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  • Mossy cloud forest occurs at higher elevations on nearly all of Borneo's mountains. These habitats are characterized by high rainfall, frequent fog, cooler temperatures, and small trees which are covered with moss and a myriad of epiphytic plants. Sarawak, Malaysia (Borneo).
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  • Rising mist spills over the forested plateau of Maliau Basin, a large conservation area in the remote interior of northern Borneo. Sabah, Malaysia (Borneo).
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  • An aerial perspective of northern Borneo's lowland rainforest reveals the heterogeneity and incredible diversity of trees that comprise this ecosystem. Various flushes of reds, oranges, and yellows among the canopy as seen here are the first indications of a periodic mass flowering event during which as many as 80% of the trees may bloom simultaneously before setting fruit. Sabah, Malaysia (Borneo).
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  • Dense mist often cloaks Borneo's rainforest at dawn, this being the water vapor rising from the transpiration of the forest itself. Sabah, Malaysia (Borneo).
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  • After several months of feeding on Aristolochia leaves, the mature caterpillar of this large birdwing butterfly (Troides andromache) prepares for its final moult into the pupal stage when metamorphosis will transform it into an adult. Sabah, Malaysia (Borneo).
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  • Although the canopy of the Borneo rainforest is bathed in the heat of the equatorial sun, as little as 5 percent of sunlight reaches the forest floor. Sabah, Malaysia (Borneo).
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  • View of canopy of lowland mixed dipterocarp forest during a mass flowering event. Sabah, Malaysia (Borneo).
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  • Reds, oranges, yellows, and other bright colors among the rainforest canopy are a sign of a mass flowering event. Borneo's rainforests undergo a cyclic mass flowering every 4-6 years (usually triggered by a dry spell), during which the majority of the trees flower and fruit simultaneously. This periodic event is a fundamental factor in the ecology of nearly all rainforest organisms in Borneo.
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  • A twisted liana winds its way into the canopy among the virgin rainforest of Tawau Hills National Park in northern Borneo. Home to some of the tallest tropical trees in the world, the forests in these hills is inhabited by a rich diversity of wildlife. Sabah, Malaysia (Borneo).
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  • Sunrise and mist over virgin rainforest in northern Borneo. Danum Valley Conservation Area, Sabah, Malaysia.
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  • Batang Ai National Park. Sarawak, Malaysia (Borneo).
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  • Sunlight filters through lowland rainforest into the entrance of a limestone cave in Mulu National Park. Sarawak, Malaysia (Borneo).
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  • Unidentified mushrooms. Sabah, Malaysia (Borneo).
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  • Scat of Mountain Treeshrew (Tupaia montana) on the peristome of Nepenthes rajah. This will be washed into the pitcher with rain where it becomes a vital source of nutrients for the plant. Sabah, Malaysia (Borneo).
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  • Mantispid (family Mantispidae), also known as a ‘mantidfly’. Looking something like a praying mantis crossed with a lacewing, this tiny predator uses the same raptorial front legs to grab its prey and hold them fast while eating them alive. Contrary to their appearance, they are not closely related to mantids, implying that this effective hunting strategy has evolved on more than one occasion in the insect kingdom. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Sunrise and mist over virgin rainforest in northern Borneo. Danum Valley Conservation Area, Sabah, Malaysia.
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  • A butterfly caterpillar (Cupha erymanthis) standing guard over a clutch of parasitic wasp pupae that recently hatched from its body. Having been impregnated by eggs from a female wasp, the larvae develop within the caterpillar, consuming its flesh as they grow. In order to keep their host alive, the wasp larvae are careful to avoid all vital organs and the caterpillar thus appears and behaves rather normal until the larvae eventually emerge. In a similar vein to the mind-controlled zombie ants, the wasp larvae also secrete some cocktail of hormones that modify the behavior of the caterpillar, in this case it is induced to stand guard and protect the wasp pupae from predators or other parasitic wasps until it eventually dies of starvation. Sarawak, Malaysia (Borneo).
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  • Morning sunlight scarcely penetrates the dense mist rising from virgin rainforest in the proposed Ulu Baleh National Park. This area is located deep in the remote interior of Sarawak, close to the geographic center of Borneo. Sarawk, Malaysia.
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  • This pond amid lowland forest is a prime breeding habitat for many species of amphibians. Sarawak, Malaysia (Borneo).
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  • Weathering from rain and wind has given rise to these razor-sharp limestone pinnacles, which can be found on the upper slopes of Mount Api in Mulu National Park. Sarawak, Malaysia (Borneo).
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  • A tannin-colored stream meanders through freshwater swamp forest in Sarawak. Trees with stilt roots, pneumatophores, and buttresses, are more abundant in this waterlogged habitat, which is frequently flooded by rains. Sarawak, Malaysia (Borneo).
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