Chien C. Lee

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  • A well-camouflaged Borneo Rainbow Toad (Ansonia latidisca) scales a mossy tree trunk in the submontane rainforest of western Borneo. Although little is known about the life history of this rare species, it is an agile climber and is probably highly arboreal. Sarawak, Malaysia (Borneo).
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  • Virgin rainforest in western Borneo. West Kalimantan, Indonesia.
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  • Distinguished by its green/yellow eyes, Hose's Bush Frog (Philautus hosii) is endemic to Borneo and considered Near Threatened due to severe habitat loss. Adults are usually found perched on vegetation over streams in lowland rainforest. Sarawak, Malaysia (Borneo).
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  • Wallace’s Flying Frog (Rhacophorus nigropalmatus) is one of the largest of all tree frogs in Borneo. It is capable of gliding down from the forest canopy by using its enlarged webbed feet as parachutes. Sarawak, Malaysia (Borneo).
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  • The Gading Flying Frog (Leptomantis gadingensis), described as recently as 2005, is a small species known from only a few localities in western Borneo.  It breeds in lowland streams. Sarawak, Malaysia (Borneo).
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  • Orchid (Dendrobium sp.). Sarawak, Malaysia (Borneo).
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  • Alocasia puncakborneensis, a new Bornean endemic described in 2020. Sarawak, Malaysia (Borneo).
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  • Long-tailed Grass Lizard (Takydromus sexlineatus). Sarawak, Malaysia (Borneo).
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  • The dainty White-eared Tree Frog (Feihyla kajau) is endemic to the rainforests of Borneo. Sarawak, Malaysia.
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  • Huntsman spider preying on camel cricket, with parasitic midge. Sarawak, Malaysia (Borneo).
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  • Orchid (Nephelaphyllum pulchrum). Sarawak, Malaysia (Borneo).
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  • Phaius subtrilobus. This large and showy terrestrial orchid is endemic to submontane forests in Borneo. Sarawak, Malaysia.
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  • A giant leaf katydid (Pseudophyllus hercules), one of the world's largest, rests in the rainforest understory. Active only at night, they use their superb camouflage to remain undetected by predators during the day. Sarawak, Malaysia (Borneo).
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  • Luminous Porecap (Favolaschia manipularis), releasing spores. Sarawak, Malaysia (Borneo).
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  • Wallace's Flying Frog (Rhacophorus nigropalmatus), male. Sarawak, Malaysia (Borneo).
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  • The tiny chlorophyll-free flowers of a Thismia (T. cf. filiformis), a mycoheterotrophic plant. Sarawak, Malaysia (Borneo).
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  • Edentistoma octosulcatum, a rare centipede that is believed to have a vegetarian diet. Sarawak, Malaysia (Borneo).
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  • Peter's Bent-toed Gecko (Cyrtodactylus consobrinus). Sarawak, Malaysia (Borneo).
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  • The bright colors on the hind wings of this tiger moth (Areas galactina, female) warn of its distasteful nature. Sarawak, Malaysia (Borneo).
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  • Cicadas (Huechys fusca), mating. Sarawak, Malaysia (Borneo).
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  • Stick insect (Dinophasma saginatum), mating. Sarawak, Malaysia (Borneo).
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  • Everett's Tree Toad (Rentapia everetti). Sarawak, Malaysia (Borneo).
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  • Dragon-headed Katydid (Lesina blanchardi), juvenile. Sarawak, Malaysia (Borneo).
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  • Mating pair of stick insects (Staelonchodes sodalis) showing extreme dimorphism between male and female of the same species. Sarawak, Malaysia (Borneo).
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  • Lowland Dwarf Toad (Pelophryne signata), male vocalising. Sarawak, Malaysia (Borneo).
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  • Darkling beetle (Strongylium sp.). Sarawak, Malaysia (Borneo).
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  • Cat Gecko (Aeluroscalabotes felinus). These slow-moving and normally extremely placid reptiles only put on an aggressive display when their other defenses (camouflage and tail dropping) have failed them. Members of the ‘eyelid geckos’, they are the only representative of the family Eublepharidae in Borneo. Sarawak, Malaysia.
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  • Katydid (Lacipoda immunda), female. Sarawak, Malaysia (Borneo).
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  • Katydid (Hapalophyllum vrazi), female. Sarawak, Malaysia (Borneo).
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  • Wallace's Flying Frog (Rhacophorus nigropalmatus), male. Sarawak, Malaysia (Borneo).
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  • Kuhl's Gliding Gecko (Gekko kuhli). Equipped with webbed feet and parasail-like flaps of skin all along its body, this tree-dwelling gecko is able to ‘parachute’ when leaping through the air to escape predators. This not only softens its landing but also enables it to glide a considerable distance. Sarawak, Malaysia (Borneo).
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  • Lanjak Bush Frog (Philautus refugii). Sarawak, Malaysia (Borneo).
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  • The White-eared Tree Frog (Feihyla kajau) lays its eggs in clusters on leaves overhanging small streams. When the tadpoles emerge they drop down into the water below. Sarawak, Malaysia (Borneo).
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  • Dwarf Toad (Pelophryne signata). Sarawak, Malaysia (Borneo).
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  • An adult male Giant Golden Orbweaver (Nephila pilipes) is dwarfed by a gigantic female of the same species.  The male's tiny size allows him to approach the agressive female unnoticed, or at least be considered nothing more than an insignificantly small prey item. Sarawak, Malaysia (Borneo).
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  • Darkling beetle (Cerogria sp.). Sarawak Malaysia (Borneo).
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  • Under the cover of darkness, a cicada sheds its nymphal skin to emerge as a fully grown adult.  Sarawak, Malaysia (Borneo).
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  • Long-legged Bush Frog (Philautus kakipanjang). Sarawak, Malaysia (Borneo).
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  • Red-legged Frog (Leptomantis rufipes). Sarawak, Malaysia (Borneo).
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  • Wallace's Flying Frog (Rhacophorus nigropalmatus). Sarawak, Malaysia (Borneo).
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  • Within the microcosm of a mossy tree stump, a tiny hidden predator lies patiently in wait. Still a juvenile, this praying mantis (Haania sp.) will eventually mature and shed most of its moss-like camouflage in favor of fully developed wings. Sarawak, Malaysia (Borneo).
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  • A recently-germinated Dipterocarp tree seedling (Shorea sp.) reaches for the sunlight on the Bornean rainforest floor.
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  • Without a single sighting in nearly 90 years, the Borneo Rainbow Toad (Ansonia latidisca) was considered possibly extinct and listed by Conservation International as one of the "10 Most Wanted Amphibians". The species was rediscovered in 2011 by a team of herpetologists from the University of Malaysia Sarawak, although it remains extremely rare and little is known of its ecology or behavior. This brings a glimmer of hope in a time of global mass amphibian extinctions. Sarawak, Malaysia (Borneo).
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  • Perhaps one of the most beautiful of all Borneo's frogs: the Borneo Flying Frog (Rhacophorus borneensis). These gliding amphibians spend most of their life in the tree canopy, only rarely descending to ground level. Sarawak, Malaysia (Borneo).
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  • Borneo Anglehead Lizard (Gonocephalus bornensis). Sarawak, Malaysia (Borneo).
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  • Borneo Necklaced Partridge (Tropicoperdix graydoni), considered by some to be a subspecies of Chestnut-necklaced Partridge (Tropicoperdix charltonii) from mainland Southeast Asia. Sabah, Malaysia (Borneo).
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  • Like other small animals of the forest understory, this Borneo Forest Dragon (Gonocephalus bornensis) must juggle his attention between searching for prey (insects and spiders) while keeping a watchful eye out for larger predators. Although this species is unable to glide like the closely-related Dracos, it is nevertheless an agile climber and can move swiftly up and down trees. The impressive mane on the back of this male signifies his maturity - we will readily defend his territory against rival males. Danum Valley, Sabah, Malaysia (Borneo).
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  • Perhaps one of the most beautiful of all Borneo's frogs: the Borneo Flying Frog (Rhacophorus borneensis). These gliding amphibians spend most of their life in the tree canopy, only rarely descending to ground level. Sarawak, Malaysia (Borneo).
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  • Spending the majority of its time underground in burrows, the Borneo Narrowmouth Toad (Gastrophrynoides borneensis) is a rarely seen species and little is known of its life history. Despite its common name and unusual appearance, this frog is a member of the Microhylidae and not a true toad. Sarawak, Malaysia (Borneo).
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  • Borneo Short-tailed Python (Python breitensteini). Sarawak, Malaysia (Borneo).
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  • Borneo Horned Frog (Pelobatrachus nasutus), female, with Matang Narrow-mouthed Frog (Microhyla nepenthicola) - comparison between one of the largest and smallest frogs in Borneo. Sarawak, Malaysia (Borneo).
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  • Borneo Horned Frog (Pelobatrachus nasutus). Sabah, Malaysia (Borneo).
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  • Without a single sighting in nearly 90 years, the Borneo Rainbow Toad (Ansonia latidisca) was considered possibly extinct and listed by Conservation International as one of the "10 Most Wanted Amphibians". The species was rediscovered in 2011 by a team of herpetologists from the University of Malaysia Sarawak, although it remains extremely rare and little is known of its ecology or behavior. This brings a glimmer of hope in a time of global mass amphibian extinctions. Sarawak, Malaysia (Borneo).
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  • Borneo Pygmy Elephant (Elephas maximus borneensis). Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Borneo Opposite-fingered Tree Frog (Feihyla inexpectata), male. Sabah, Malaysia (Borneo).
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  • Unseen since 1924, the Borneo Rainbow Toad (Ansonia latidisca) was previously listed as one of the "world's top 10 most wanted frogs" by Conservation International in their Global Search for Lost Frogs in 2010. It was rediscovered in Sarawak in 2011. Sarawak, Malaysia (Borneo).
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  • Dense mist often cloaks Borneo's rainforest at dawn, this being the water vapor rising from the transpiration of the forest itself. Sabah, Malaysia (Borneo).
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  • A twisted liana winds its way into the canopy among the virgin rainforest of Tawau Hills National Park in northern Borneo. Home to some of the tallest tropical trees in the world, the forests in these hills is inhabited by a rich diversity of wildlife. Sabah, Malaysia (Borneo).
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  • Borneo Frog-eating Snake (Stegonotus borneensis), juvenile. Sarawak, Malaysia (Borneo).
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  • Although the canopy of the Borneo rainforest is bathed in the heat of the equatorial sun, as little as 5 percent of sunlight reaches the forest floor. Sabah, Malaysia (Borneo).
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  • Reds, oranges, yellows, and other bright colors among the rainforest canopy are a sign of a mass flowering event. Borneo's rainforests undergo a cyclic mass flowering every 4-6 years (usually triggered by a dry spell), during which the majority of the trees flower and fruit simultaneously. This periodic event is a fundamental factor in the ecology of nearly all rainforest organisms in Borneo.
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  • After several months of feeding on Aristolochia leaves, the mature caterpillar of this large birdwing butterfly (Troides andromache) prepares for its final moult into the pupal stage when metamorphosis will transform it into an adult. Sabah, Malaysia (Borneo).
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  • View of canopy of lowland mixed dipterocarp forest during a mass flowering event. Sabah, Malaysia (Borneo).
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  • An aerial perspective of northern Borneo's lowland rainforest reveals the heterogeneity and incredible diversity of trees that comprise this ecosystem. Various flushes of reds, oranges, and yellows among the canopy as seen here are the first indications of a periodic mass flowering event during which as many as 80% of the trees may bloom simultaneously before setting fruit. Sabah, Malaysia (Borneo).
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  • Borneo Forest Dragon (Gonocephalus bornensis). Sabah, Malaysia (Borneo).
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  • Heavy rain falls over a small river amid lowland rainforest. Many inland regions of Borneo such as this receive as much as 4 meters of rainfall per year. Sarawak, Malaysia (Borneo).
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  • Mossy cloud forest occurs at higher elevations on nearly all of Borneo's mountains. These habitats are characterized by high rainfall, frequent fog, cooler temperatures, and small trees which are covered with moss and a myriad of epiphytic plants. Sarawak, Malaysia (Borneo).
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  • Rising mist spills over the forested plateau of Maliau Basin, a large conservation area in the remote interior of northern Borneo. Sabah, Malaysia (Borneo).
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  • Sunrise and mist over virgin rainforest in northern Borneo. Danum Valley Conservation Area, Sabah, Malaysia.
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  • Batang Ai National Park. Sarawak, Malaysia (Borneo).
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  • Oil Palm (Elaeis guineensis) plantation showing small riparian buffer zones along rivers. Northern Borneo.
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  • Scat of Mountain Treeshrew (Tupaia montana) on the peristome of Nepenthes rajah. This will be washed into the pitcher with rain where it becomes a vital source of nutrients for the plant. Sabah, Malaysia (Borneo).
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  • Sunrise and mist over virgin rainforest in northern Borneo. Danum Valley Conservation Area, Sabah, Malaysia.
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  • Sometimes referred to as 'durian kura-kura' (tortoise durian), Durio testudinarum is one of the rarest of edible durian species. It flowers near the base of the tree and produces clusters of small fruits that bear a musky odor. Like other durians, the blooms of D. testudinarum are open at night, though the pollinator of this species remains unknown. Sarawak, Malaysia (Borneo).
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  • A tannin-colored stream meanders through freshwater swamp forest in Sarawak. Trees with stilt roots, pneumatophores, and buttresses, are more abundant in this waterlogged habitat, which is frequently flooded by rains. Sarawak, Malaysia (Borneo).
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  • An aerial view reveals the thick mist rising from the canopy after heavy rains on virgin rainforest in northern Borneo. These forests often receive over 4000ml of rain annually. Sabah, Malaysia.
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  • Sunrise and mist over virgin rainforest in northern Borneo. Danum Valley Conservation Area, Sabah, Malaysia.
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  • Large stretches of virgin rainforest still remain in the remote interior of eastern Borneo. East Kalimantan, Indonesia.
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  • Oil Palm (Elaeis guineensis) plantation showing small riparian buffer zones along rivers. Northern Borneo.
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  • A butterfly caterpillar (Cupha erymanthis) standing guard over a clutch of parasitic wasp pupae that recently hatched from its body. Having been impregnated by eggs from a female wasp, the larvae develop within the caterpillar, consuming its flesh as they grow. In order to keep their host alive, the wasp larvae are careful to avoid all vital organs and the caterpillar thus appears and behaves rather normal until the larvae eventually emerge. In a similar vein to the mind-controlled zombie ants, the wasp larvae also secrete some cocktail of hormones that modify the behavior of the caterpillar, in this case it is induced to stand guard and protect the wasp pupae from predators or other parasitic wasps until it eventually dies of starvation. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Weathering from rain and wind has given rise to these razor-sharp limestone pinnacles, which can be found on the upper slopes of Mount Api in Mulu National Park. Sarawak, Malaysia (Borneo).
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  • Sunlight filters through lowland rainforest into the entrance of a limestone cave in Mulu National Park. Sarawak, Malaysia (Borneo).
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  • Large granite boulders decorate the coastline at Tanjung Datu National Park at the western tip of Borneo. Sarawak, Malaysia.
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  • Unidentified mushrooms. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Setting sun and thundershowers over the rugged landscape of West Kalimantan, Borneo.
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  • The beautiful 3-tiered Takob-Akob Waterfall is located deep within a steep sandstone gorge of Maliau Basin. Sabah, Malaysia (Borneo).
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  • An aerial view of virgin rainforest. Sabah, Malaysia (Borneo).
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  • Morning sunlight scarcely penetrates the dense mist rising from virgin rainforest in the proposed Ulu Baleh National Park. This area is located deep in the remote interior of Sarawak, close to the geographic center of Borneo. Sarawk, Malaysia.
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  • Crystal clear streams flow from undisturbed rainforest habitats in the interior of Batang Ai National Park in southern Sarawak, Malaysia (Borneo).
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  • Mantispid (family Mantispidae), also known as a ‘mantidfly’. Looking something like a praying mantis crossed with a lacewing, this tiny predator uses the same raptorial front legs to grab its prey and hold them fast while eating them alive. Contrary to their appearance, they are not closely related to mantids, implying that this effective hunting strategy has evolved on more than one occasion in the insect kingdom. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • This pond amid lowland forest is a prime breeding habitat for many species of amphibians. Sarawak, Malaysia (Borneo).
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  • Large sandstone blocks litter a hillside at Payeh Maga in northern Borneo. Sarawak, Malaysia.
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  • Large sandstone boulders and stunted elfin forest the summit plateau of Gunung Murud, Sarawak's highest mountain. Sarawak, Malaysia (Borneo).
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  • One of the many waterfalls draining the remote highland Payeh Maga plateau. Sarawak, Malaysia (Borneo).
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  • An increasingly rare sight, clear rocky streams are an important and indicative component of pristine habitats in Borneo rainforests. Sarawak, Malaysia.
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  • Undoubtedly one of the rarest and most peculiar of all Borneo's reptiles: the Bornean Horned Lizard (Harpesaurus borneensis). The males of this species are unique in possessing a distinctive horn at the tip of their snout, a trait not seen in the females nor found in any other Bornean lizard. Even more unusual, this is the only known agamid lizard in Southeast Asia to give birth to live young rather than laying eggs. As only a few specimens of this species have ever been found, its full life history and distribution in Borneo are still largely unknown. Sarawak, Malaysia (Borneo).
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  • Nycticebus menagensis. One of four species of Slow Loris found in Borneo. All Slow Loris are nocturnal arboreal primates which feed on small animals, insects and soft fruits. Sarawak, Malaysia (Borneo).
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  • In the Borneo rainforest, leaves are not always what they seem. Here, a beam of sunlight filtering down from the canopy illuminates an extraordinary insect. Officially described and named just last year, this leaf katydid (Eulophophyllum lobulatum) is one of two species in its genus on mainland Borneo that share the unusual wings and leaf-like legs, rendering it a marvel of animal crypsis. Although this male specimen is lime-green, females bear a striking pinkish coloration. Mount Kinabalu, Sabah, Malaysia (Borneo).
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  • The Black-bearded Tomb Bat (Taphozous melanopogon) roosts in caves and large rock crevices, emerging at night to feed on insects high above the forest canopy. It is found throughout much of mainland Southeast Asia as well as Borneo, Sumatra, and Java. It's echolocation calls while flying are audible as high pitched clicks. Sarawak, Malaysia (Borneo).
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  • Endemic to the mountains of northern Borneo, the Kinabalu Horned Frog (Pelobatrachus baluensis) breeds only in clear rocky streams with cold water. By day they lie camouflaged among the leaf litter on the forest floor, but they emerge at night to call for mates. Sabah, Malaysia (Borneo).
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