Chien C. Lee

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  • Triangle Keelback (Xenochrophis trianguligerus). Maliau Basin Conservation Area, Sabah, Malaysia (Borneo).
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  • Black-headed Cat Snake (Boiga nigriceps). Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Although usually considered a terrestrial species, the Striped Kukri Snake (Oligodon octolineatus) is an excellent climber and can occasionally be seen climbing tree trunks. Sarawak, Malaysia (Borneo).
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  • Brown Tree Snake (Boiga irregularis), juvenile. Mamberamo Basin, Papua, Indonesia (New Guinea).
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  • Paradise Tree Snake (Chrysopelea paradisi). Sarawak, Malaysia (Borneo).
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  • Spotted Cat Snake (Boiga drapiezii). Sarawak, Malaysia (Borneo).
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  • The Fire-lipped Keelback (Rhabdophis murudensis) is a rare snake known only from the higher mountains of northern Borneo.  When threatened, it has the remarkable ability to exude a white milky venom from the back of its neck as a defensive mechanism. Gunung Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • Malayan Banded Wolf Snake (Lycodon subcinctus), juvenile. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Brown-tailed Racer (Gonyosoma oxycephalum). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • White-spotted Cat Snake (Boiga drapiezii), green morph. Sarawak, Malaysia (Borneo).
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  • The Twin-barred Flying Snake (Chrysopelea pelias) is the smallest and rarest of Southeast Asia's flying snake species. Like other Chrysopelea, this snake hunts lizards and is almost completely arboreal. Sarawak, Malaysia (Borneo).
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  • Bronzeback (Dendrelaphis terrificus). Gorontalo, Sulawesi, Indonesia.
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  • A juvenile Speckle-bellied Keelback (Rhabdophis chrysargos) from the forested slopes of Mount Tambuyukon in northern Borneo. This natricine snake has a mildly toxic saliva and feeds primarily on frogs. Sabah, Malaysia (Borneo).
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  • Barnes' Cat Snake (Boiga barnesii). Sinharaja National Park, Sri Lanka.
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  • Cloudy Snail-eating Snake (Sibon nebulatus). Limón, Costa Rica.
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  • Catesby's Snail-eater (Dipsas catesbyi). Yasuní National Park, Ecuador.
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  • Black-headed Cat Snake (Boiga nigriceps), juvenile. Kubah National Park, Sarawak, Malaysia (Borneo).
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  • Cloudy Snail-eating Snake (Sibon nebulatus). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • Mangrove Cat Snake (Boiga dendrophila). Sabah, Malaysia (Borneo).
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  • Red-headed Reed Snake (Calamaria schlegeli schlegeli), a small soil-dwelling species which feeds on frogs and slugs. Sarawak, Malaysia (Borneo).
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  • Blunthead Tree Snake (Imantodes cenchoa), a lizard hunting specialist that due to its extremely slender body is able to seek out sleeping lizards on even the most delicate of branches. Yasuní National Park, Ecuador.
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  • Paradise Tree Snake (Chrysopelea paradisi). Sarawak, Malaysia (Borneo).
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  • Wolf snake (Stegonotus sp.). Papua, Indonesia (New Guinea).
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  • The Oriental Vine Snake (Ahaetulla prasina) is an agile climber, feeding on small birds and lizards with its excellent binocular vision. Sabah, Malaysia (Borneo).
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  • The Mangrove Cat Snake (Boiga dendrophila) is a widespread inhabitant of coastal and riverine forests in Southeast Asia, where it hunts for small mammals, reptiles (including other snakes), and birds. A rear-fanged snake, research has revealed that its venom is specifically tailored to affect birds and is relatively ineffective in subduing mammals (although bites in humans can cause pain and inflammation). This is the first known occurrence of a bird-specific toxin. Sarawak, Malaysia (Borneo).
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  • The rare Forsten's Pointed Snake (Rabdion forsteni) is endemic to Sulawesi and known from only a few specimens. South Sulawesi, Indonesia.
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  • Paradise Flying Snake (Chrysopelea paradisi). Sarawak, Malaysia (Borneo).
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  • Yellow-striped Ratsnake (Coelognathus flavolineatus). Kampot, Cambodia.
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  • A Harlequin Gliding Frog (Rhacophorus pardalis) struggles futilely as it is being slowly swallowed alive by a Black-Headed Cat Snake (Boiga nigriceps). Stagnant pools of water in the Borneo rainforest serve as important breeding grounds for many tree frogs which gather around in great numbers, a fact that some predators wisely take advantage of. Kubah National Park, Sarawak, Malaysia (Borneo).
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  • A juvenile Oriental Vine Snake (Ahaetulla prasina) basks in a beam of sunlight in the rainforest of Tangkoko National Park. These highly arboreal snakes are excellent climbers and are adept at hunting lizards, their favored prey. North Sulawesi, Indonesia.
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  • Three-banded Bridled Snake (Lycodon tristrigatus). Sarawak, Malaysia (Borneo).
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  • The Black-headed Cat Snake (Boiga nigriceps) is an arboreal predator of birds and other snakes. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Sabah Keelback (Hebius flavifrons). Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • The Beauty Ratsnake (Elaphe taeniura grabowskyi) is an adept climber, capable of scaling limestone cave walls where it feeds on roosting bats and swiftlets. Sarawak, Malaysia (Borneo).
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  • Neotropical Snail-Eater (Dipsas indica). Yasuní National Park, Ecuador.
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  • Venezuela’s fabled tabletop mountains may have once inspired Arthur Conan Doyle’s science fiction novel “The Lost World”, where he imagined the summits to be home to a plethora of gigantic prehistoric creatures that had been separated from the evolution of life on the rest of the planet for eons of time. Disappointingly, dinosaurs were never found on these mountains and, despite being home to many endemics and indeed a fair share of “living fossils”, recent genetic studies have shown that many species in fact have likely arrived well after the tepuis had been fully formed. This reveals that the imposing rocky cliffs, which seemingly isolate the summits from the jungles below, may not be as significant a physical barrier for dispersal as once believed. This young Boddaert's Tropical Racer (Mastigodryas boddaerti), encountered on the plateau of Auyán Tepui, is a widespread species in northern South America, and although prey is scarce on these mountaintops, it presumably hunts for endemic frogs.  Canaima National Park, Venezuela.
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  • Striped Bronzeback (Dendrelaphis caudolineatus). Sarawak, Malaysia (Borneo).
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  • Striped Sharpnose Snake (Philodryas argentea). Yasuní National Park, Ecuador.
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  • Elegant Snail-Eater (Dipsas elegans). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • Borneo Frog-eating Snake (Stegonotus borneensis), juvenile. Sarawak, Malaysia (Borneo).
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  • Keel-bellied Vine Snake (Dryophiops rubescens), a lizard-eating arboreal colubrid. Sarawak, Malaysia (Borneo).
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  • Olive Mountain Keelback (Opisthotropis typica). Sarawak, Malaysia (Borneo).
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  • Although usually considered a terrestrial species, the Striped Kukri Snake (Oligodon octolineatus) is an excellent climber and can occasionally be seen climbing tree trunks. Sarawak, Malaysia (Borneo).
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  • One of the most beautiful of the Boigas: the Jasper Cat Snake (Boiga jaspidea), a scarce species in Borneo. Like other Boigas, it is a rear-fanged snake and hunts primarily in the trees. Sarawak, Malaysia (Borneo).
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  • Rough-backed Litter Snake (Xenodermus javanicus). Sarawak, Malaysia (Borneo).
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  • Dryad Snake (Mastigodryas melanolomus). Osa Peninsula, Puntarenas, Costa Rica.
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  • Black-headed Cat Snake (Boiga nigriceps). Ulu Temburong National Park, Brunei (Borneo).
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  • Spotted Cat Snake (Boiga drapiezii). Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • The Black-headed Cat Snake (Boiga nigriceps) is an arboreal predator of birds and other snakes.  Although it has one of the most potent venoms of all colubrid snakes, it possesses only rear fangs and is not an aggressive species. Sarawak, Malaysia (Borneo).
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  • Madagascar Cat-eyed Snake (Madagascarophis colubrinus). Mahajanga, Madagascar.
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  • The Three-striped Ground Snake (Pseudoxyrhopus tritaeniatus) is certainly one of the most beautiful of Madagascan endemic snakes, but why is it colored this way? Bold red stripes are frequently used as an aposematic warning, indicating to other animals of severe danger. It is curious then that not only is this species completely non-venomous but there are in fact no other dangerous snakes in Madagascar that this could possibly be a mimic of. An interesting theory points to the co-occurrence of large red toxic millipedes in the same forests – could these actually be a mimicry model for this snake? On an island legendary for its biological curiosities, nothing seems beyond the realm of possibility. Ranomafana National Park, Madagascar.
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  • Malagasy Giant Hognose Snake (Leioheterodon madagascariensis), hunting for lizard eggs which are buried in the sand. Akanin'ny Nofy, Madagascar.
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  • Common Big-eyed Snake (Mimophis mahfalensis). Toliara, Madagascar.
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  • Madagascar Cat-eyed Snake (Madagascarophis colubrinus). Antsiranana, Madagascar.
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  • Forest Water Snake (Thamnosophis infrasignatus). Ranomafana National Park, Madagascar.
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  • Malagasy Giant Hognose Snake (Leioheterodon madagascariensis), a burrowing species and Madagascar's largest colubrid. Tomasina, Madagascar.
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  • Sometimes referred to as the ‘Trees of Life’, baobabs (Adansonia spp.) provide food and shelter for many species of animals, particularly those surviving in the brutally hot and arid climate of Madagascar’s southwest. Here, a Southwestern Night Snake (Ithycyphus oursi) takes shelter during the heat of the day beneath a fissure in the bark of a Za Baobab (A. za). These hiding spots are sometimes shared with geckos, scorpions and Madagascar’s famous hissing cockroaches. This snake, incidentally, is one of the few venomous species on the island; while perhaps not dangerous to humans its bite has been observed to cause paralysis in chameleons. Amboasary, Madagascar.
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