Chien C. Lee

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  • Mountain Treeshrew (Tupaia montana) feeding at Nepenthes rajah.  Recent research has shown the the world's largest pitcher plant Nepenthes rajah is not exclusively carnivorous. Like the related N. lowii, this species attracts treeshrews by secreting nectar on the undersurface of the lid. These animals frequently leave their droppings in the pitcher, which serves as a valuable nitrogen source in their impoverished mountain habitat. Mount Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • An infrared camera trap reveals a secretive nocturnal visitor to the giant pitchers of Nepenthes rajah. The Kinabalu Rat (Rattus baluensis) is known only from the upper slopes of Mount Kinabalu in northern Borneo. New research has confirmed that this rodent shares a similar mutualistic relationship with these pitcher plants as the Mountain Treeshrew (Tupaia montana), by obtaining sweet nectar in exchange for their nitrogen-rich droppings. Sabah, Malaysia (Borneo).
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  • Kayan Loris (Nycticebus kayan), juvenile. Gunung Mulu National Park, Sarawak, Malaysia (Borneo).
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  • A Sunda Pangolin (Manis javanica) forages at night for ant and termite nests, using its keen sense of smell to locate a nest. Sarawak, Malaysia (Borneo).
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  • The rigid armour-like scales of the Pangolin (Manis javanica) afford it a high degree of protection from predators.  The scales are considered as valuable in Chinese medicine, and the animal is frequently hunted for this reason. Sarawak, Malaysia (Borneo).
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  • An uncommon species in Sarawak, the Sunda Free-tailed Bat (Mops mops) often forages at night high above forest clearings for insects. It often roosts in hollow trees. Batang Ai National Park, Sarawak, Malaysia (Borneo).
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  • Slender Mongoose (Herpestes sanguineus). Mpumalanga, South Africa.
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  • Proboscis Monkey (Nasalis larvatus). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Sri Lankan Grey Langur (Semnopithecus priam thersites). Yala National Park, Sri Lanka.
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  • Nycticebus menagensis. One of four species of Slow Loris found in Borneo. All Slow Loris are nocturnal arboreal primates which feed on small animals, insects and soft fruits. Sarawak, Malaysia (Borneo).
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  • The Black-bearded Tomb Bat (Taphozous melanopogon) roosts in caves and large rock crevices, emerging at night to feed on insects high above the forest canopy. It is found throughout much of mainland Southeast Asia as well as Borneo, Sumatra, and Java. It's echolocation calls while flying are audible as high pitched clicks. Sarawak, Malaysia (Borneo).
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  • Threatened by habitat loss and hunting pressure, the Crested Black Macaque (Macaca nigra nigra) is now listed as Critically Endangered. This species lives only in forested areas at the northernmost tip of the island of Sulawesi. North Sulawesi, Indonesia.
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  • Spectacled Flying Fox (Pteropus conspicillatus). Waigeo Island, West Papua, Indonesia.
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  • Smallest of all the anteaters, the Silky Anteater (Cyclopes didactylus) is seldom seen because it spends much of its life high in the rainforest canopy, often curled up as an indistinguishable ball of fluff. They have no teeth and can only defend themselves by means of their razor sharp sickle-like fore-claws, which are usually used for tearing open ant nests. Females bear a single youngster at a time, which is carried on their back until large enough to feed on its own. Esmeraldas, Ecuador.
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  • The Long-tongued Nectar Bat (Macroglossus minimus) is an important pollinator of many rainforest trees. Here it is feeding on the nectar of banana flowers. Sarawak, Malaysia (Borneo).
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  • Polynesian Rat (Rattus exulans). Sarawak, Malaysia (Borneo).
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  • Spotted Cuscus (Spilocuscus maculatus chrysorrhous), juvenile. Papua, Indonesia (New Guinea).
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  • Owston's Palm Civet (Chrotogale owstoni), a nocturnal carnivore from Laos and Vietnam, is a highly vulnerable species due to illegal poaching. Captive. Endangered Primate Rescue Centre, Cuc Phuong National Park, Vietnam.
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  • Lesser Bare-backed fruit Bat (Dobsonia minor), feeding on fig. Nimbokrang, Papua, Indonesia (New Guinea).
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  • With a mouthful of dry moss, a Whitehead's Pygmy Squirrel (Exilisciurus whiteheadi) pauses cautiously to make sure nobody is watching before disappearing into its nest in a rock crevice. Endemic to the island of Borneo, this tiny squirrel lives only in cool montane rainforests.
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  • Goodman's Mouse Lemur (Microcebus lehilahytsara), feeding on the nectar of night-blooming Dombeya flowers. Andasibe, Madagascar.
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  • Sri Lanka Giant Squirrel (Ratufa macroura dandolena). Central Province, Sri Lanka.
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  • A Short-nosed Fruit Bat (Cynopterus brachyotis) swoops in to select a ripe fig (Ficus fistulosa) growing directly on the trunk of the tree. Rather than eating on the spot, the bat will carry the fig off to a favorite perch where it can dine in safety, thus helping to spread the tree's seeds. Sarawak, Malaysia (Borneo).
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  • Black-and-white Ruffed Lemur (Varecia variegata), a Critically Endangered species and undoubtedly one of the most beautiful of all lemurs. Entirely dependent on tall undisturbed rainforests, these lemurs are threatened by habitat loss and are suffering from an island-wide population decline. Ranomafana National Park, Madagascar.
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  • Bornean Orangutan (Pongo pygmaeus), infant. Sarawak, Malaysia (Borneo).
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  • By means of its extensive skin membranes, the Sunda Colugo (Galeopterus variegatus borneanus), is capable of gliding impressive distances between trees. This adult is carrying a young offspring on its underside. Bearing no relation to true lemurs, colugos are in their own unique order and are only distantly related to primates and treeshrews. They are nocturnal, feeding on leaves, flowers, and sap. Recent research on morphological and genetic variations indicate that the Bornean and Javan subspecies may merit recognition as distinct species. Sarawak, Malaysia (Borneo).
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  • Marañón White-fronted Capuchin (Cebus yuracus). Yasuní National Park, Ecuador.
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  • Hose’s Civet (Diplogale hosei), one of Borneo’s rarest and least known endemic mammals. This otter-like carnivore is an inhabitant of high elevation mossy forests where it hunts for frogs, fish, and other small animals. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Endemic to the Bird's Head Peninsula of western New Guinea, the Vogelkop Tree Kangaroo (Dendrolagus ursinus) has become increasingly threatened by habitat loss and hunting. West Papua, Indonesia (New Guinea).
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  • One of Borneo's most famous endemic mammals, the Proboscis Monkey (Nasalis larvatus) is restricted to coastal swamp forests and mangroves. Bako National Park, Sarawak, Malaysia (Borneo).
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  • Nearly a splitting image of its namesake, a Lesser Hedgehog Tenrec (Echinops telfari) is a remarkable example of convergent evolution. All of Madagascar's tenrecs, which are more closely related to golden moles and elephant shrews, are descendent from a single colonizing ancestor, which has diversified into a variety of species to fill various niches. This particularly spiny one is native to the island's arid southwest, where it forages for ground insects much like the European hedgehogs. Toliara, Madagascar.
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  • Deep within spiral of an unfurled Ravenala leaf, a Madagascar Sucker-footed Bat (Myzopoda aurita) roosts in safety. With its adhesive pads it is able to cling to the smooth surface of the leaf where even insects cannot find a foothold. In this way, Myzopoda are remarkably free of the ectoparasites which plague most other bat species. Fianarantsoa, Madagascar.
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  • Madagascar’s largest land predator, the Fossa (Cryptoprocta ferox). This receptive female has climbed a tall tree where she waits for a male who is able to join her on one of the upper branches. Reaching her isn’t always an easy task for the males, which are considerably larger and heavier, and they often face fierce competition when attempting to do so. Kirindy Reserve, Madagascar.
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  • The Long-tongued Nectar Bat (Macroglossus minimus) is an important pollinator of many rainforest trees including mangroves and bananas. Sarawak, Malaysia (Borneo).
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  • Fresh tracks of an Oriental Small-clawed Otter (Aonyx cinerea) along the beach at Tanjung Datu National Park. Sarawak, Malaysia (Borneo).
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  • Whitehead's Pigmy Squirrel (Exilisciurus whiteheadi), also known as the Tufted Pigmy Squirrel, is endemic to the mountains of Borneo.  It feeds almost entirely on the mosses and lichens growing on tree trunks. Sabah, Malaysia (Borneo).
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  • Lowland Streaked Tenrec (Hemicentetes semispinosus). Like a miniature porcupine, these indomitable tiny mammals protect themselves from predators by an armament of barbed spines. They feed on insects and other invertebrates which they find in the leaf litter of the forest floor using their long sensitive snout. All species of tenrec are endemic to Madagascar. Mantadia National Park, Madagascar.
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  • Lion (Panthera leo). Moremi Game Reserve, Ngamiland, Botswana.
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  • The small and nocturnal Sunda Stink-badger (Mydaus javanensis lucifer) is a relative of the skunks (family Mephitidae) occurring in Borneo, Sumatra, and Java. It feeds on invertebrates and small animals as well as tuberous roots. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • A large male Bornean Clouded Leopard (Neofelis diardi) is caught on camera trap as he patrols his territory in the rainforest of Danum Valley, Sabah. Borneo's largest cat species, the clouded leopard preys on a variety of smaller wildlife ranging from deer and wild boar to monkeys. Sabah, Malaysia (Borneo).
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  • The least known of all of Sulawesi's monkeys, Heck's Macaque (Macaca hecki) occurs on the northwestern portion of the island. Five other endemic species of macaque are distributed in different regions of Sulawesi, occasionally producing hybrids where their ranges overlap. All are diurnal and feed largely on fruits. Central Sulawesi, Indonesia.
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  • The nocturnal blossoms of a wild durian (Durio kutejensis) are visited by a Cave Nectar Bat (Eonycteris spelaea). Although also attracting bees, birds, and other small pollinators during the day, the flowers of this tree open fully at dusk whereupon they begin to produce copious amounts of rich mango-scented nectar – an enticing draw for a bat with a keen sense of smell and an irresistible sweet tooth. With their ability to travel great distances and visit countless flowers in a single night, nectar-feeding bats play a vital role in the pollination of many rainforest plants. Sarawak, Malaysia (Borneo).
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  • Mareeba Rock Wallaby (Petrogale mareeba). Queensland, Australia.
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  • Hills covered with Traveller's Palm (Ravenala madagascariensis), habitat for Madagascar Sucker-footed Bat (Myzopoda aurita) which roost in the young unfurled leaves. Fianarantsoa, Madagascar.
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  • Napo Saki (Pithecia napensis). Yasuní National Park, Ecuador.
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  • Scat of Mountain Treeshrew (Tupaia montana) on the peristome of Nepenthes rajah. This will be washed into the pitcher with rain where it becomes a vital source of nutrients for the plant. Sabah, Malaysia (Borneo).
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  • The least known of all of Sulawesi's monkeys, Heck's Macaque occurs on the northwestern portion of the island. Five other endemic species of macaque are distributed in different regions of Sulawesi, occasionally producing hybrids where their ranges overlap. All are diurnal and feed largely on fruits. Central Sulawesi, Indonesia.
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  • Sunda Colugo (Galeopterus variegatus), adult female with young. Sarawak, Malaysia (Borneo).
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  • Verreaux's Sifaka (Propithecus verreauxi), crossing road. Popularly known as the ‘Dancing Sifakas’ for their remarkable hopping agility, few visitors realize that the dancing spectacle is in part due to the loss of these animals’ habitat: they naturally dwell in dry and spiny forests and only descend to the ground when they must cross a cleared area. Amboasary, Madagascar.
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  • The bright orange coloration of this infant Silvered Langur (Trachypithecus cristatus) helps them to be seen and hence guarded by the entire troop. They will gradually lose this coloration after 3 months of age. Bako National Park, Sarawak, Malaysia (Borneo).
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  • Smallest of all the anteaters, the Silky Anteater (Cyclopes didactylus) is seldom seen because it spends much of its life high in the rainforest canopy, often curled up as an indistinguishable ball of fluff. They have no teeth and can only defend themselves by means of their razor sharp sickle-like fore-claws, which are usually used for tearing open ant nests. Females bear a single youngster at a time, which is carried on their back until large enough to feed on its own. Esmeraldas, Ecuador.
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  • Sugar Glider (Petaurus breviceps), feeding on sap of Acacia tree. Wasur National Park, Papua, Indonesia (New Guinea).
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  • Vogelkop Ringtail (Pseudochirulus schlegeli), one of New Guinea’s least studied possums, being known from only a few specimens. Like other ringtails, they are strictly nocturnal, emerging at night to feed on fruit and leaves. Arfak Mountains, West Papua, Indonesia (New Guinea).
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  • Gursky's Spectral Tarsier (Tarsius spectrumgurskyae), juvenile. North Sulawesi, Indonesia.
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  • The endemic Sulawesi Warty Pig (Sus celebensis) occurs in a variety of habitats ranging from lowland to montane rainforest. Although its only natural predator is the Reticulated Python (Python reticulatus), its populations across the island have been in decline due to habitat loss. Gorontalo, Indonesia.
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  • Nectar-feeding bats play an essential role in the pollination of many rainforest trees. Here, a Long-tongued Nectar Bat (Macroglossus minimus) arrives at the night-blooming blossoms of a durian tree (Durio zibethinus). Prized for its delectable aromatic fruit, the productivity of many durian plantations is under threat where the native bat populations have been killed off by hunters. Sabah, Malaysia (Borneo).
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  • Silvered Leaf Monkey (Trachypithecus cristatus). Sarawak, Malaysia (Borneo).
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  • A pair of Lesser False Vampire Bats (Megaderma spasma) roosting near the entrance of a shallow limestone cave. Named because of their superficial resemblance to true vampire bats (which are restricted to Central and South America), Megaderma are insectivorous and do not drink blood. This species sometimes attains a large size (nearly 35g in weight) and is known to occasionally take more substantial prey including lizards, small birds, mammals, and even other bats. Sarawak, Malaysia (Borneo).
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  • Short-nosed Fruit Bats (Cynopterus brachyotis) roosting under a wild banana leaf. Sabah, Malaysia (Borneo).
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  • The Crested Black Macaque (Macaca nigra nigra) is one of four monkey species endemic to Sulawesi.  They are sometimes wrongly referred to as apes because their tail is extremely small and difficult to see.  They travel mostly on the ground in large groups of up to nearly 100 individuals. North Sulawesi, Indonesia.
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  • After quickly selecting a ripe fig (Ficus fistulosa), a Short-nosed Fruit Bat (Cynopterus brachyotis) makes off with her prize to dine some distance away at a safe perch, thereby helping to disperse the tree's seeds that will ultimately lead to fruit for her future generations. Figs reach their pinnacle of diversity in Borneo with at least 150 species and, perhaps more than any other group of plants, are considered keystone species of the rainforest because of the complex interdependences they exhibit with countless animals and insects. Sarawak, Malaysia (Borneo).
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  • The Maroon Langur (Presbytis rubicunda) is endemic to Borneo and nearby Karimata Island, with several subspecies occurring throughout its range. Like other leaf monkeys they are almost completely arboreal and feed on foliage, seeds, and some fruits.  Danum Valley Conservation Area, Sabah, Malaysia.
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  • The Smooth Otter (Lutrogale perspicillata) is the largest otter species in Southeast Asia, reaching up to 1.3 meters in length. Smooth Otters mate for life and hunt cooperatively for fish. Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Owston's Palm Civet (Chrotogale owstoni), a nocturnal carnivore from Laos and Vietnam, is a highly vulnerable species due to illegal poaching. Captive. Endangered Primate Rescue Centre, Cuc Phuong National Park, Vietnam.
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  • A Trefoil Horseshoe Bat (Rhinolophus trifoliatus) rests while hanging from a twig of an understory forest tree in Danum Valley. Sabah, Malaysia (Borneo).
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  • This large male Northwest Bornean Orangutan (Pongo pygmaeus pygmaeus) is just beginning to develop cheek pads, a sign of maturity and placing his age to perhaps between 15 to 20 years. Batang Ai, Sarawak, Malaysia (Borneo).
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  • A Small-toothed Sportive Lemur (Lepilemur microdon) peers curiously from its tree hole in the montane forest of Ranomafana National Park, Madagascar. Usually emerging only at night, sportive lemurs frequently wake to keep a watchful eye out for intruders into their territory. Not long ago only 8 species of Lepilemur were known from Madagascar, but recent molecular work has resulted in at least 26 distinct species now being recognized, with more sure to follow.
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  • A Bornean Bearded Pig (Sus barbatus) forages for food in a mangrove swamp during low tide.
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  • A female Bornean Orang-utan (Pongo pygmaeus) with her young infant. Semengoh Forest Reserve, Sarawak, Malaysia (Borneo).
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  • The newly distinguished Kayan Loris (Nycticebus kayan) differs from other Bornean loris species by a combination of morphological traits. Sarawak, Malaysia (Borneo).
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  • Feared by some, revered by others, the Aye-aye (Daubentonia madagascariensis) is surely one of Madagascar's most bizarre lemurs. Unique among primates, the Aye-aye's front incisors grow continuously like a rodent, leading to the early belief that they were some sort of gigantic nocturnal squirrel. Equally unusual are its long bony middle fingers which are used by the animal to tap tree trunks for the sound of insect grubs inside and then extract them like a fish hook. Unfortunately, Aye-ayes have become gravely endangered from habitat loss and persecution by people: they are often killed on sight due to the superstition that they are an omen of death. Tomasina, Madagascar.
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  • Deer Cave (located in Mulu National Park) is one of the world's largest cave passages, with an opening chamber over 120 meters in height. It is home to an estimated 5 million bats. Sarawak, Malaysia (Borneo).
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  • Impala (Aepyceros melampus). Kruger National Park, Mpumalanga, South Africa.
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  • The Western Tarsier (Cephalopachus bancanus borneanus), also known as Horsfield's Tarsier, is a small nocturnal primate occurring in the lowland forests of Sumatra and Borneo. They are carnivorous and feed mainly on insects, but occasionally take birds and bats. Sarawak, Malaysia (Borneo).
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  • The rare and enigmatic Babirusa (Babyrousa celebensis) occurs only on Sulawesi and a few neighboring islands. Adult males have four conspicuous tusks which are formed from their elongated canine teeth. Gorontalo, Sulawesi, Indonesia.
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  • A Gursky's Spectral Tarsier (Tarsius spectrumgurskyae) perches outside of its tree hole roost. These tiny nocturnal primates emerge every evening to hunt for insects. Tangkoko Nature Reserve, North Sulawesi, Indonesia.
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  • A Horsfield's Tarsier (Cephalopachus bancanus) on the prowl for food in the rainforest of western Borneo. These tiny primates are exclusively carnivorous, hunting insects, small lizards, and even snakes by leaping deftly through the thick forest understory and snatching prey with their long nimble fingers. Unlike most other nocturnal mammals which possess a reflective layer behind their retina to help with night vision, tarsiers instead compensate by having enormous eyes. In fact, this species sports the largest eyes in relation to its body size of any mammal in the world - these are locked in place and the animal is only able to shift its vision by rotating its head. Sarawak, Malaysia (Borneo).
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  • The Sumatran Surili (Presbytis melalophos) is endemic to the island of Sumatra.  Like other leaf monkeys, this species is primarily folivorous, but also consumes flowers and fruits. Kerinci Seblat National Park, West Sumatra, Indonesia.
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  • The critically endangered Greater Bamboo Lemur (Prolemur simus) is restricted to wet rainforests of Madagascar's southeast where it is highly dependent on Cathariostachys bamboo. Ranomafana National Park, Madagascar.
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  • With fewer than a thousand individuals remaining in the wild, the Critically Endangered Golden Bamboo Lemur (Hapalemur aureus) holds on to a tenuous existence in the rainforests of Madagascar. It’s discovery in the mid-1980’s was a crucial factor that led to the foundation of the now famous Ranomafana National Park – a reserve that protects over 400 sq. km. of tropical forest. Like other bamboo lemurs, it feeds primarily on bamboo shoots which, due to their high cyanide content, are lethally toxic to most other animals. Ranomafana National Park, Madagascar.
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  • Spectacled Flying Fox (Pteropus conspicillatus). Waigeo Island, West Papua, Indonesia.
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  • Large Flying Fox (Pteropus vampyrus). Sarawak, Malaysia (Borneo).
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  • A Red-shanked Douc (Pygathrix nemaeus) munches on a handful of young vine leaves it has picked in the forest canopy. Although these endangered monkeys feed on over sixty different species of plants, they select their food carefully and in particular avoid sweet fruits which can upset the balance of bacteria in their guts. Like other leaf-eating colobine monkeys, doucs have large chambered stomachs – making them the only ruminant primates and giving them their characteristic pot-bellied appearance. Son Tra Nature Reserve, Vietnam.
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  • A Maroon Langur (Presbytis rubicunda) pauses for a break while munching on his chosen leaf. Endemic to Borneo and a few offshore islands, these leaf monkeys feed on a wide range of foliage and unripe fruits, traveling in groups of up to a dozen individuals. One theory regarding their distinctive bright orange coloration is that this may help to confuse some predators into mistaking them for a much larger and stronger Orangutan, and thus deter attacks. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • With fewer than a thousand individuals remaining in the wild, the Critically Endangered Golden Bamboo Lemur (Hapalemur aureus) holds on to a tenuous existence in the rainforests of Madagascar. It’s discovery in the mid-1980’s was a crucial factor that led to the foundation of the now famous Ranomafana National Park – a reserve that protects over 400 sq. km. of tropical forest. Like other bamboo lemurs, it feeds primarily on bamboo shoots which, due to their high cyanide content, are lethally toxic to most other animals. Ranomafana National Park, Madagascar.
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  • Sulawesi Babirusa (Babyrousa celebensis). Gorontalo, Sulawesi.
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  • Ceylon Elephant (Elephas maximus maximus). Yala National Park, Sri Lanka.
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  • Northeast Bornean Orangutan (Pongo pygmaeus morio), adult male. Sabah, Malaysia (Borneo).
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  • One of the world's most endangered primates, the Bornean Banded Langur (Presbytis chrysomelas chrysomelas) has been threatened by both habitat loss and hunting. Now, only a few isolated populations remain in coastal Sarawak, with an estimated total of just a few hundred individuals. They feed on foliage and fruits in lowland rainforest. Sarawak, Malaysia (Borneo).
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  • Although the blooms of this rare mangrove tree (Pelliciera rhizophorae) are visited by hummingbirds in the day, the flowers remain open at night and exude a strong musty odor to attract an entirely different animal. Captured by an infrared camera trigger, a Leaf-nosed Bat (Glossophaga soricina) approaches an open flower to partake of the rich source of nectar. Nectar-feeding bats are common in the tropics and are responsible for the pollination of a great number of tree species. Choco, Colombia.
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  • Deep within spiral of an unfurled Ravenala leaf, a Madagascar Sucker-footed Bat (Myzopoda aurita) roosts in safety. With its adhesive pads it is able to cling to the smooth surface of the leaf where even insects cannot find a foothold. In this way, Myzopoda are remarkably free of the ectoparasites which plague most other bat species. Fianarantsoa, Madagascar.
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  • Intermediate Roundleaf Bat (Hipposideros larvatus). Yunnan, China.
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  • Grey Mouse Lemur (Microcebus murinus). Kirindy Reserve, Madagascar.
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  • A large colony of Wrinkle-lipped Bats (Chaerephon plicata) emerges from a cave at dusk. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • The Brown Spiny Rat (Maxomys rajah) is found in lowland rainforests where it forages for food on the ground and in low vegetation. Sarawak, Malaysia (Borneo).
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  • The Dusky Fruit Bat (Penthetor lucasi) roosts in large colonies in caves, though like most other fruit bats it is not capable of echolocation and relies on its keen eyesight for navigation.  It emerges at night to feed on a variety of fruits, often carrying food back to its roost to eat. Sarawak, Malaysia (Borneo).
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  • A newly distinguished species, the Kayan Loris (Nycticebus kayan) differs from other Bornean loris species by a combination of morphological traits. Sarawak, Malaysia (Borneo).
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  • A view of a hanging Trefoil Horseshoe Bat (Rhinolophus trifoliatus) from below reveals its remarkable face. The elaborate noseleaves help the bat to focus its echolocation calls, with the various parts shaped for different frequencies. Sabah, Malaysia (Borneo).
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  • Although shy, the Bearded Pig (Sus barbatus) is abundant in the forests of Borneo, and its presence is usually indicated by diggings and wallows left as they forage for food on the forest floor. Sarawak, Malaysia (Borneo).
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  • An adult male Silvered Leaf Monkey (Trachypithecus cristatus) keeps a watchful eye out for any threats to his harem of females. Sarawak, Malaysia (Borneo).
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