Chien C. Lee

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  • Black-lipped Shrub Lizard (Pelturagonia nigrilabris). Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Flat-nosed Pitviper (Craspedocephalus puniceus). West Java, Indonesia.
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  • Triangle Keelback (Xenochrophis trianguligerus). Maliau Basin Conservation Area, Sabah, Malaysia (Borneo).
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  • One of the world’s least studied crocodilians, the False Gharial (Tomistoma schlegelii) is a rare inhabitant of the dark tannin colored waters of peat swamps in Borneo, Sumatra, and Malaysia. Due to their narrow snout they were once believed to feed exclusively on fish, however large specimens can reach lengths of over 5 meters and have more recently been observed to consume prey as large as Proboscis Monkeys, or in one unfortunate instance, a grown man. Although much remains unknown about their life history, it is clear that they are rapidly becoming endangered due to destruction of their peat swamp habitats. Sarawak, Malaysia (Borneo).
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  • Kuhl's Gliding Gecko (Gekko kuhli). By means of its large webbed feet and lateral skin flaps, this arboreal gecko is capable of gliding or parachuting between trees. Sarawak, Malaysia (Borneo).
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  • Kuhl's Gliding Gecko (Gekko kuhli). Equipped with webbed feet and parasail-like flaps of skin all along its body, this tree-dwelling gecko is able to ‘parachute’ when leaping through the air to escape predators. This not only softens its landing but also enables it to glide a considerable distance. Sarawak, Malaysia (Borneo).
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  • The rare Kinabalu Brown Pit Viper (Garthius chaseni) is known only from a few mountains in northern Borneo where it occurs in high elevation forest. Very little is known of its life history as it has seldom been observed in the wild. Sabah, Malaysia (Borneo).
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  • Large-headed Shrub Lizard (Pelturagonia cephalum). Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • Black-lipped Shrub Lizard (Pelturagonia nigrilabris). Sabah, Malaysia (Borneo).
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  • The superbly camouflaged Kinabalu Gliding Gecko (Gekko rhacophorus) is endemic to the montane forests of Mount Kinabalu in northern Borneo.  This poorly-known species is a nocturnal insectivorous tree-dweller and, like other geckos of this genus, possesses skin flaps to assist in a parachuting flight. Sabah, Malaysia (Borneo).
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  • Still wet from a passing rainstorm, a Sabah Bamboo Pitviper (Trimeresurus sabahi) lies in ambush on the side of a tree, waiting for the approach of suitable prey. By maintaining a low metabolism and sometimes remaining in a single position for weeks at a time, tropical vipers such as this can survive by feeding as infrequently as once every few months. Mount Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • White-spotted Cat Snake (Boiga drapiezii), green morph. Sarawak, Malaysia (Borneo).
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  • Few creatures possess a mastery of camouflage the way Madagascar’s leaf-tailed geckos do. This Mossy Leaf-tailed Gecko (Uroplatus sikorae) sleeps by day on a rainforest liana, its outline obscured by frilled dermal flaps that line its body. By always resting in a downward-facing position they may be able to quickly catch unsuspecting insects that climb up from the ground and also conceal their reflective lidless eyes from predators lurking above. Ranomafana National Park, Madagascar.
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  • Madagascar Giant Day Gecko (Phelsuma grandis). Canal des Pangalanes, Madagascar.
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  • Cryptic Chameleon (Calumma crypticum), male. Ranomafana National Park, Madagascar.
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  • Ecuadorian Toadheaded Pitviper (Bothrocophias campbelli). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • The Twin-barred Flying Snake (Chrysopelea pelias) is the smallest and rarest of Southeast Asia's flying snake species. Like other Chrysopelea, this snake hunts lizards and is almost completely arboreal. Sarawak, Malaysia (Borneo).
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  • Malayan Flat-shelled Turtle (Notochelys platynota). Sarawak, Malaysia (Borneo).
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  • Sabah Bamboo Pitviper (Trimeresurus sabahi). Sarawak, Malaysia (Borneo).
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  • Although the Banded Coral Snake (Calliophis intestinalis) is not agressive, it possesses an extremely toxic venom like other Elapids. When disturbed it displays its red tail and the brightly colored bands on its undersurface which serve as a warning signal to would-be predators. Sarawak, Malaysia (Borneo).
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  • Flat-nosed Pitviper (Craspedocephalus puniceus). West Java, Indonesia.
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  • Blue-eyed Angle-headed Lizard (Gonocephalus liogaster), male. Sarawak, Malaysia (Borneo).
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  • A juvenile Parson's Chameleon (Calumma parsoni). More frequently occurring in shades of green, this is one of the world's largest chameleons, with adult specimens sometimes reaching the size of a house cat. They are endemic to rainforest habitats on Madagascar and are threatened by both habitat loss and collection for the international pet trade. Andasibe, Madagascar.
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  • Bronzeback (Dendrelaphis terrificus). Gorontalo, Sulawesi, Indonesia.
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  • Spiny-tailed Gecko (Diplodactylus ciliaris). Exmouth, Western Australia.
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  • When threatened, the Knob-tailed Gecko (Nephrurus levis) raises its body off the ground to appear larger in size. Cape Range National Park, Western Australia.
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  • Hikida's Forest Dragon (Hypsilurus hikidanus). Papua, Indonesia (New Guinea).
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  • The mimicry of a dead leaf is so precise for the Satanic Leaf-tailed Gecko (Uroplatus phantasticus), they can often hide effectively in plain sight. It has been untested to what degree these geckos actively select a substrate that matches their individual patterns, but U. phantasticus appears to typically hide among dead foliage which suits it perfectly. Ranomafana National Park, Madagascar.
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  • Long-tailed Grass Lizard (Takydromus sexlineatus). Sarawak, Malaysia (Borneo).
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  • Paradise Tree Snake (Chrysopelea paradisi). Sarawak, Malaysia (Borneo).
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  • A study on perfection in mimicry: a Satanic Leaf-tailed Gecko (Uroplatus phantasticus). One of 14 species of leaf-tailed geckos endemic to Madagascar, this nocturnal insectivore utilizes its incredible camouflage to hide among dead foliage by day. All Uroplatus geckos are under severe threat from deforestation and over-collecting for the international pet trade. Andasibe, Madagascar.
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  • The Bornean Keeled Green Pit Viper (Tropidolaemus subannulatus) is an arboreal species, often laying in wait for prey on the same branch for weeks at a time. Sarawak, Malaysia (Borneo).
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  • Bornean Short Python (Python breitensteini). Sarawak, Malaysia (Borneo).
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  • Common Sun Skink (Eutropis multifasciata). Sarawak, Malaysia (Borneo).
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  • Black-headed Cat Snake (Boiga nigriceps). Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Four-clawed Gecko (Gehyra mutilata) attending a lanternfly (Pyrops whiteheadi) for secreted honeydew. Host tree is Pometia pinnata. Sabah, Malaysia (Borneo).
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  • Purple Anole (Anolis purpurescens). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • Southeast Asian Softshell Turtle (Amyda cartilaginea). Sepilok Forest Reserve, Sabah, Malaysia (Borneo).
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  • Vogel’s Pit Viper (Trimeresurus vogeli). Bach Ma National Park, Vietnam.
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  • Although usually considered a terrestrial species, the Striped Kukri Snake (Oligodon octolineatus) is an excellent climber and can occasionally be seen climbing tree trunks. Sarawak, Malaysia (Borneo).
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  • Males of the Madagascar's Satanic Leaf-tailed Gecko (Uroplatus phantasticus) can usually be distinguished by their elaborately notched tails, a trait which gives them better mimicry of a leaf that has been partially eaten by insects. Exactly why this is a sexually dimorphic feature in these geckos is unknown, but in some animals different camouflage patterns in males and females can arise where each sex spends their time hiding in different locations and thus face differing selective pressures on their appearance. Ranomafana National Park, Madagascar.
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  • Sumatran Pit Viper (Trimeresurus sumatranus). Sarawak, Malaysia (Borneo).
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  • Blotch Bow-finger Gecko (Cyrtodactylus yakhuna). North Western Province, Sri Lanka.
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  • Hump-nosed lizard photo (Lyriocephalus scutatus), male. Sinharaja National Park, Southern Province, Sri Lanka.
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  • Barnes' Cat Snake (Boiga barnesii). Sinharaja National Park, Sri Lanka.
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  • Horned Flying Lizard (Draco cornutus), male using his dewlap to display for courtship and territorial aggression. Sabah, Malaysia (Borneo).
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  • Undoubtedly one of the rarest and most peculiar of all Borneo's reptiles: the Bornean Horned Lizard (Harpesaurus borneensis). The males of this species are unique in possessing a distinctive horn at the tip of their snout, a trait not seen in the females nor found in any other Bornean lizard. Even more unusual, this is the only known agamid lizard in Southeast Asia to give birth to live young rather than laying eggs. As only a few specimens of this species have ever been found, its full life history and distribution in Borneo are still largely unknown. Sarawak, Malaysia (Borneo).
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  • While pythons undoubtedly hold the crown for the largest and most famous constricting snakes of Indonesia, a single genus of boas also occurs in the rainforests of the eastern archipelago. Often overlooked, the Pacific Keel-scaled Boa (Candoia carinata) seldom exceeds a meter in length. It is an ambush predator of frogs and lizards, and like other boas, bears live young rather than laying eggs. Digul River, Papua, Indonesia (New Guinea).
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  • Borneo Anglehead Lizard (Gonocephalus bornensis). Sarawak, Malaysia (Borneo).
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  • Spotted Cat Snake (Boiga drapiezii). Sarawak, Malaysia (Borneo).
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  • The largest terrestrial and river predator in the world, the Estuarine Crocodile (Crocodylus porosus) can reach lengths of over 6 m and consumes a wide range of animal prey. They are commonly found in mangroves, lagoons, and large rivers along the coastline of Borneo.  Kinabatangan River, Sabah, Malaysia (Borneo).
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  • Bornean Angle-headed Lizard (Gonocephalus bornensis), male. Sarawak, Malaysia (Borneo).
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  • Paradise Tree Snake (Chrysopelea paradisi). Sarawak, Malaysia (Borneo).
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  • Lined Pipe Snake (Cylindrophis lineatus) also known as Blanford's Pipe Snake, a rare species endemic to Borneo and known from only a few specimens. When threatened, this species defends itself by raising its tail which is thickened and colored to appear as a head. Sarawak, Malaysia (Borneo).
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  • The Fire-lipped Keelback (Rhabdophis murudensis) is a rare snake known only from the higher mountains of northern Borneo.  When threatened, it has the remarkable ability to exude a white milky venom from the back of its neck as a defensive mechanism. Gunung Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • A species of bent-toed gecko (Cyrtodactylus cf. mikianus) from the Mamberamo Basin in western New Guinea. Papua, Indonesia.
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  • A master of camouflage, the Mossy Leaf-tailed Gecko (Uroplatus sikorae) is almost completely undetectable when it is at rest on a small tree in the forest understory. Frilled flaps of skin along its body break up its outline, making it appear nearly seamless with the tree on which it rests. Andasibe, Madagascar.
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  • A juvenile Oriental Vine Snake (Ahaetulla prasina) basks in a beam of sunlight in the rainforest of Tangkoko National Park. These highly arboreal snakes are excellent climbers and are adept at hunting lizards, their favored prey. North Sulawesi, Indonesia.
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  • A juvenile Rock Monitor (Varanus albigularis). Mpumalanga, South Africa.
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  • The impressive Sailfin Lizard (Hydrosaurus celebensis) is the largest agamid lizard in the world, reaching over 1 m in length.  They are very agile swimmers and are seldom found far from rivers.  They feed on leaves and insects. Central Sulawesi, Indonesia.
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  • A large male O'Shaughnessy's Chameleon (Calumma oshaughnessyi) stalks up a forest liana, keeping an eye out for both prey and predator alike. With 360-degree vision, remarkable image acuity, and monocular depth perception, chameleons can detect insects that might be well out of reach, judge their distance accurately, and even distinguishing them from their surroundings when the insect is completely still. Ranomafana National Park, Madagascar.
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  • Spiny Ground Gecko (Paroedura bastardi). Amboasary, Madagascar.
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  • Warty Chameleon (Furcifer verrucosus). Toliara, Madagascar.
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  • This female Madagascar Pygmy Kingfisher (Corythornis madagascariensis) has just been gifted a small lizard (Madascincus melanopleura) by her mate as part of a courtship ritual. Despite their name, these birds are not dependent on water, and typically hunt for insects and small vertebrates on the forest floor. Ranomafana National Park, Madagascar.
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  • Sap-feeding bugs such as this lanternfly (Pyrops whiteheadi) ingest large quantities of the nutrient-poor fluid, excreting the excess in the form of honeydew. Here a gecko (Gehyra mutilata) waits below the insect for an opportunistic meal of the sweet fluid. Sabah, Malaysia (Borneo).
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  • Red-tailed Pipe Snake (Cylindrophis ruffus). Sarawak, Malaysia (Borneo).
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  • A juvenile Speckle-bellied Keelback (Rhabdophis chrysargos) from the forested slopes of Mount Tambuyukon in northern Borneo. This natricine snake has a mildly toxic saliva and feeds primarily on frogs. Sabah, Malaysia (Borneo).
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  • Amazon Tree Boa (Corallus hortulanus). Yasuní National Park, Ecuador.
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  • Malayan Banded Wolf Snake (Lycodon subcinctus), juvenile. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Brown-tailed Racer (Gonyosoma oxycephalum). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Like other small animals of the forest understory, this Borneo Forest Dragon (Gonocephalus bornensis) must juggle his attention between searching for prey (insects and spiders) while keeping a watchful eye out for larger predators. Although this species is unable to glide like the closely-related Dracos, it is nevertheless an agile climber and can move swiftly up and down trees. The impressive mane on the back of this male signifies his maturity - we will readily defend his territory against rival males. Danum Valley, Sabah, Malaysia (Borneo).
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  • Despite its appearance and usually fearsome disposition, the Dusky Mock Viper (Psammodynastes pulverulentus) possesses only a weak venom and is not dangerous to humans. It feeds on lizards and frogs. Sabah, Malaysia (Borneo).
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  • The Three-striped Ground Snake (Pseudoxyrhopus tritaeniatus) is certainly one of the most beautiful of Madagascan endemic snakes, but why is it colored this way? Bold red stripes are frequently used as an aposematic warning, indicating to other animals of severe danger. It is curious then that not only is this species completely non-venomous but there are in fact no other dangerous snakes in Madagascar that this could possibly be a mimic of. An interesting theory points to the co-occurrence of large red toxic millipedes in the same forests – could these actually be a mimicry model for this snake? On an island legendary for its biological curiosities, nothing seems beyond the realm of possibility. Ranomafana National Park, Madagascar.
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  • Forest Water Snake (Thamnosophis infrasignatus). Ranomafana National Park, Madagascar.
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  • Mossy Leaf-tailed Gecko (Uroplatus sikorae). Mantadia National Park, Madagascar.
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  • Cloudy Snail-eating Snake (Sibon nebulatus). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • Hump-nosed Viper (Hypnale zara). Sinharaja National Park, Sri Lanka.
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  • Endemic to the mountains of central Sri Lanka, the Rhino Horn Lizard (Ceratophora stoddartii) is named for the unusual spine found on the snout of males. This is likely a product of female mate selection. Central Province, Sri Lanka.
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  • Brown Tree Snake (Boiga irregularis), juvenile. Mamberamo Basin, Papua, Indonesia (New Guinea).
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  • Boulenger's Green Anole (Anolis chloris), male displaying. Choco, Colombia.
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  • The bold coloration of the Red-headed Krait (Bungarus flaviceps) is a warning to would-be predators of the highly potent venom it possesses. This large elapid feeds primarily on other reptiles including lizards and snakes. Sarawak, Malaysia (Borneo).
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  • Mocquard's Madagascar Ground Gecko (Paroedura bastardi), juvenile. Tsingy de Bemaraha National Park, Madagascar.
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  • Although completely non-venomous, the Painted Mock Viper (Psammodynastes pictus) defends itself by aggressively striking if threatened. It often hunts aquatic animals such as frogs and fish by perching on a low branch over the surface of the water. Sarawak, Malaysia (Borneo).
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  • The Earless Monitor Lizard (Lanthanotus borneensis) is endemic to Borneo where it lives in rocky stream habitats in lowland rainforest. Because of its extreme rarity (it is known from only a few specimens), its life history remains unstudied. It is the only species within its entire family (Lanthanotidae) and was once though to be the missing link between lizards and snakes. Sarawak, Malaysia (Borneo).
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  • A Harlequin Gliding Frog (Rhacophorus pardalis) struggles futilely as it is being slowly swallowed alive by a Black-Headed Cat Snake (Boiga nigriceps). Stagnant pools of water in the Borneo rainforest serve as important breeding grounds for many tree frogs which gather around in great numbers, a fact that some predators wisely take advantage of. Kubah National Park, Sarawak, Malaysia (Borneo).
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  • Horsfield's Gliding Gecko (Gekko horsfieldii). Sabah, Malaysia (Borneo).
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  • Peter's Bent-toed Gecko (Cyrtodactylus consobrinus). Sarawak, Malaysia (Borneo).
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  • The Oriental Vine Snake (Ahaetulla prasina) is an agile climber, feeding on small birds and lizards with its excellent binocular vision. Sabah, Malaysia (Borneo).
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  • Two-banded Chameleon (Furcifer balteatus), female. Ranomafana National Park, Madagascar.
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  • Bornean Keeled Green Pit Viper (Tropidolaemus subannulatus). Sarawak, Malaysia (Borneo).
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  • A Pilbara Death Adder (Acanthophis wellsi) from the desert regions of Western Australia. Although superficially resembling vipers by their stout body and habit of ambushing prey, death adders are actually elapids and more closely related to cobras, kraits, and coral snakes. Cape Range National Park, Western Australia.
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  • A young Parson's Chameleon (Calumma parsonii) in the rainforest of Masoala National Park. Antsiranana, Madagascar.
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  • Venezuela’s fabled tabletop mountains may have once inspired Arthur Conan Doyle’s science fiction novel “The Lost World”, where he imagined the summits to be home to a plethora of gigantic prehistoric creatures that had been separated from the evolution of life on the rest of the planet for eons of time. Disappointingly, dinosaurs were never found on these mountains and, despite being home to many endemics and indeed a fair share of “living fossils”, recent genetic studies have shown that many species in fact have likely arrived well after the tepuis had been fully formed. This reveals that the imposing rocky cliffs, which seemingly isolate the summits from the jungles below, may not be as significant a physical barrier for dispersal as once believed. This young Boddaert's Tropical Racer (Mastigodryas boddaerti), encountered on the plateau of Auyán Tepui, is a widespread species in northern South America, and although prey is scarce on these mountaintops, it presumably hunts for endemic frogs.  Canaima National Park, Venezuela.
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  • Green Tree Python (Morelia azurea), juvenile.
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  • Banded Malaysian Coralsnake (Calliophis intestinalis). Sarawak, Malaysia (Borneo).
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  • Cloudy Snail-eating Snake (Sibon nebulatus). Limón, Costa Rica.
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  • Catesby's Snail-eater (Dipsas catesbyi). Yasuní National Park, Ecuador.
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  • Southern New Guinea Crocodile (Crocodylus halli), juvenile. Wasur National Park, Papua, Indonesia (New Guinea).
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  • Three-banded Bridled Snake (Lycodon tristrigatus). Sarawak, Malaysia (Borneo).
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  • Asian Brown Tortoise (Manouria emys), male. Sabah, Malaysia (Borneo).
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