Chien C. Lee

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  • Sabah Bamboo Pitviper (Trimeresurus sabahi). Sarawak, Malaysia (Borneo).
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  • Flat-nosed Pitviper (Craspedocephalus puniceus). West Java, Indonesia.
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  • Black-lipped Shrub Lizard (Pelturagonia nigrilabris). Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Flat-nosed Pitviper (Craspedocephalus puniceus). West Java, Indonesia.
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  • Horsfield's Gliding Gecko (Gekko horsfieldii). Sabah, Malaysia (Borneo).
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  • Kuhl's Gliding Gecko (Gekko kuhli). By means of its large webbed feet and lateral skin flaps, this arboreal gecko is capable of gliding or parachuting between trees. Sarawak, Malaysia (Borneo).
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  • Kuhl's Gliding Gecko (Gekko kuhli). Equipped with webbed feet and parasail-like flaps of skin all along its body, this tree-dwelling gecko is able to ‘parachute’ when leaping through the air to escape predators. This not only softens its landing but also enables it to glide a considerable distance. Sarawak, Malaysia (Borneo).
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  • Triangle Keelback (Xenochrophis trianguligerus). Maliau Basin Conservation Area, Sabah, Malaysia (Borneo).
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  • The Bornean Leaf-nosed Pit Viper (Craspedocephalus borneensis) is a well-camouflaged snake, frequently occuring on the forest floor or in low vegetation. They strike readily when disturbed. Sarawak, Malaysia (Borneo).
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  • The superbly camouflaged Kinabalu Gliding Gecko (Gekko rhacophorus) is endemic to the montane forests of Mount Kinabalu in northern Borneo.  This poorly-known species is a nocturnal insectivorous tree-dweller and, like other geckos of this genus, possesses skin flaps to assist in a parachuting flight. Sabah, Malaysia (Borneo).
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  • The rare Kinabalu Brown Pit Viper (Garthius chaseni) is known only from a few mountains in northern Borneo where it occurs in high elevation forest. Very little is known of its life history as it has seldom been observed in the wild. Sabah, Malaysia (Borneo).
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  • The Spiny-headed Shrub Lizard (Pelturagonia spiniceps) is known only from montane forest at a few localities in northwestern Borneo. Pulong Tau National Park, Sarawak, Malaysia (Borneo).
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  • Black-lipped Shrub Lizard (Pelturagonia nigrilabris). Sabah, Malaysia (Borneo).
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  • Still wet from a passing rainstorm, a Sabah Bamboo Pitviper (Trimeresurus sabahi) lies in ambush on the side of a tree, waiting for the approach of suitable prey. By maintaining a low metabolism and sometimes remaining in a single position for weeks at a time, tropical vipers such as this can survive by feeding as infrequently as once every few months. Mount Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • White-spotted Cat Snake (Boiga drapiezii), green morph. Sarawak, Malaysia (Borneo).
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  • The Twin-barred Flying Snake (Chrysopelea pelias) is the smallest and rarest of Southeast Asia's flying snake species. Like other Chrysopelea, this snake hunts lizards and is almost completely arboreal. Sarawak, Malaysia (Borneo).
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  • Malayan Flat-shelled Turtle (Notochelys platynota). Sarawak, Malaysia (Borneo).
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  • Bronzeback (Dendrelaphis terrificus). Gorontalo, Sulawesi, Indonesia.
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  • When threatened, the Knob-tailed Gecko (Nephrurus levis) raises its body off the ground to appear larger in size. Cape Range National Park, Western Australia.
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  • Hikida's Forest Dragon (Hypsilurus hikidanus). Central Papua, Indonesia (New Guinea).
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  • Madagascar Giant Day Gecko (Phelsuma grandis). Canal des Pangalanes, Madagascar.
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  • Cryptic Chameleon (Calumma crypticum), male. Ranomafana National Park, Madagascar.
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  • Although the Banded Coral Snake (Calliophis intestinalis) is not aggressive, it possesses an extremely toxic venom like other Elapids. When disturbed it displays its red tail and the brightly colored bands on its undersurface which serve as a warning signal to would-be predators. Sarawak, Malaysia (Borneo).
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  • Blue-eyed Angle-headed Lizard (Gonocephalus liogaster), male. Sarawak, Malaysia (Borneo).
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  • A juvenile Parson's Chameleon (Calumma parsoni). More frequently occurring in shades of green, this is one of the world's largest chameleons, with adult specimens sometimes reaching the size of a house cat. They are endemic to rainforest habitats on Madagascar and are threatened by both habitat loss and collection for the international pet trade. Andasibe, Madagascar.
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  • Spiny-tailed Gecko (Diplodactylus ciliaris). Exmouth, Western Australia.
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  • The mimicry of a dead leaf is so precise for the Satanic Leaf-tailed Gecko (Uroplatus phantasticus), they can often hide effectively in plain sight. It has been untested to what degree these geckos actively select a substrate that matches their individual patterns, but U. phantasticus appears to typically hide among dead foliage which suits it perfectly. Ranomafana National Park, Madagascar.
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  • Long-tailed Grass Lizard (Takydromus sexlineatus). Sarawak, Malaysia (Borneo).
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  • Paradise Tree Snake (Chrysopelea paradisi). Sarawak, Malaysia (Borneo).
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  • A study on perfection in mimicry: a Satanic Leaf-tailed Gecko (Uroplatus phantasticus). One of 14 species of leaf-tailed geckos endemic to Madagascar, this nocturnal insectivore utilizes its incredible camouflage to hide among dead foliage by day. All Uroplatus geckos are under severe threat from deforestation and over-collecting for the international pet trade. Andasibe, Madagascar.
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  • The Bornean Keeled Green Pit Viper (Tropidolaemus subannulatus) is an arboreal species, often laying in wait for prey on the same branch for weeks at a time. Sarawak, Malaysia (Borneo).
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  • Sharp-nosed Chameleon (Kinyongia oxyrhina), male. Udzungwa Mountains, Tanzania.
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  • Dumeril's Monitor (Varanus dumerilii), juvenile. Sarawak, Malaysia (Borneo).
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  • A large male O'Shaughnessy's Chameleon (Calumma oshaughnessyi) stalks up a forest liana, keeping an eye out for both prey and predator alike. With 360-degree vision, remarkable image acuity, and monocular depth perception, chameleons can detect insects that might be well out of reach, judge their distance accurately, and even distinguishing them from their surroundings when the insect is completely still. Ranomafana National Park, Madagascar.
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  • Common Sun Skink (Eutropis multifasciata). Sarawak, Malaysia (Borneo).
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  • Spiny Ground Gecko (Paroedura bastardi). Amboasary, Madagascar.
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  • This female Madagascar Pygmy Kingfisher (Corythornis madagascariensis) has just been gifted a small lizard (Madascincus melanopleura) by her mate as part of a courtship ritual. Despite their name, these birds are not dependent on water, and typically hunt for insects and small vertebrates on the forest floor. Ranomafana National Park, Madagascar.
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  • Red-tailed Pipe Snake (Cylindrophis ruffus). Sarawak, Malaysia (Borneo).
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  • Four-clawed Gecko (Gehyra mutilata) attending a lanternfly (Pyrops whiteheadi) for secreted honeydew. Host tree is Pometia pinnata. Sabah, Malaysia (Borneo).
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  • A juvenile Speckle-bellied Keelback (Rhabdophis chrysargos) from the forested slopes of Mount Tambuyukon in northern Borneo. This natricine snake has a mildly toxic saliva and feeds primarily on frogs. Sabah, Malaysia (Borneo).
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  • Southeast Asian Softshell Turtle (Amyda cartilaginea). Sepilok Forest Reserve, Sabah, Malaysia (Borneo).
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  • Amazon Tree Boa (Corallus hortulanus). Yasuní National Park, Ecuador.
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  • Malayan Banded Wolf Snake (Lycodon subcinctus), juvenile. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Brown-tailed Racer (Gonyosoma oxycephalum). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Like other small animals of the forest understory, this Borneo Forest Dragon (Gonocephalus bornensis) must juggle his attention between searching for prey (insects and spiders) while keeping a watchful eye out for larger predators. Although this species is unable to glide like the closely-related Dracos, it is nevertheless an agile climber and can move swiftly up and down trees. The impressive mane on the back of this male signifies his maturity - we will readily defend his territory against rival males. Danum Valley, Sabah, Malaysia (Borneo).
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  • Despite its appearance and usually fearsome disposition, the Dusky Mock Viper (Psammodynastes pulverulentus) possesses only a weak venom and is not dangerous to humans. It feeds on lizards and frogs. Sabah, Malaysia (Borneo).
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  • The Three-striped Ground Snake (Pseudoxyrhopus tritaeniatus) is certainly one of the most beautiful of Madagascan endemic snakes, but why is it colored this way? Bold red stripes are frequently used as an aposematic warning, indicating to other animals of severe danger. It is curious then that not only is this species completely non-venomous but there are in fact no other dangerous snakes in Madagascar that this could possibly be a mimic of. An interesting theory points to the co-occurrence of large red toxic millipedes in the same forests – could these actually be a mimicry model for this snake? On an island legendary for its biological curiosities, nothing seems beyond the realm of possibility. Ranomafana National Park, Madagascar.
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  • Kirindy Leaf-toed Gecko (Paroedura rennerae), juvenile. Tsingy de Bemaraha National Park, Madagascar.
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  • The Earless Monitor Lizard (Lanthanotus borneensis) is endemic to Borneo where it lives in rocky stream habitats in lowland rainforest. Because of its extreme rarity (it is known from only a few specimens), its life history remains unstudied. It is the only species within its entire family (Lanthanotidae) and was once though to be the missing link between lizards and snakes. Sarawak, Malaysia (Borneo).
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  • A Harlequin Gliding Frog (Rhacophorus pardalis) struggles futilely as it is being slowly swallowed alive by a Black-Headed Cat Snake (Boiga nigriceps). Stagnant pools of water in the Borneo rainforest serve as important breeding grounds for many tree frogs which gather around in great numbers, a fact that some predators wisely take advantage of. Kubah National Park, Sarawak, Malaysia (Borneo).
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  • Two-banded Chameleon (Furcifer balteatus), female. Ranomafana National Park, Madagascar.
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  • Bornean Keeled Green Pit Viper (Tropidolaemus subannulatus). Sarawak, Malaysia (Borneo).
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  • A species of bent-toed gecko (Cyrtodactylus cf. mikianus) from the Mamberamo Basin in western New Guinea. Papua, Indonesia.
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  • A juvenile Oriental Vine Snake (Ahaetulla prasina) basks in a beam of sunlight in the rainforest of Tangkoko National Park. These highly arboreal snakes are excellent climbers and are adept at hunting lizards, their favored prey. North Sulawesi, Indonesia.
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  • A juvenile Rock Monitor (Varanus albigularis). Mpumalanga, South Africa.
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  • Canopy Slug-eating Snake (Sibon canopy), a new species described in January 2023, endemic to Panama. Panamá Oeste, Panama.
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  • Usambara Eyelash Viper (Atheris ceratophora). Udzungwa Mountains, Tanzania.
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  • Bornean Short Python (Python breitensteini). Sarawak, Malaysia (Borneo).
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  • Black-headed Cat Snake (Boiga nigriceps). Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Purple Anole (Anolis purpurescens). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • Scarcely longer than an earthworm, this juvenile slug-eating snake (Pareas hamptoni) sleeps balanced on top of a fern frond. Many slug-eating snakes retain the  unusual asymmetric dentition of their snail-eating ancestors, an adaptation for pulling snails out of their shells via a process known as ‘mandibular walking’. Bach Ma National Park, Vietnam.
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  • Sumatran Pit Viper (Trimeresurus sumatranus). Sarawak, Malaysia (Borneo).
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  • Mossy Leaf-tailed Gecko (Uroplatus sikorae). Mantadia National Park, Madagascar.
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  • Vieira's Snail-eating Snake (Sibon vieirai). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • Horned Flying Lizard (Draco cornutus), male using his dewlap to display for courtship and territorial aggression. Sabah, Malaysia (Borneo).
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  • Red-headed Reed Snake (Calamaria schlegeli schlegeli), a small soil-dwelling species which feeds on frogs and slugs. Sarawak, Malaysia (Borneo).
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  • Blunthead Tree Snake (Imantodes cenchoa), a lizard hunting specialist that due to its extremely slender body is able to seek out sleeping lizards on even the most delicate of branches. Yasuní National Park, Ecuador.
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  • The bold coloration of the Red-headed Krait (Bungarus flaviceps) is a warning to would-be predators of the highly potent venom it possesses. This large elapid feeds primarily on other reptiles including lizards and snakes. Sarawak, Malaysia (Borneo).
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  • Although masterful visual predators by day, chameleons lack light-sensitive rods in their eyes and thus have poor night vision. Because of this they typically retire at dusk to a sleeping perch, often returning to a favorite exposed branch which ideally puts them out of the reach of nocturnal snakes. This dozing Two-banded Chameleon (Furcifer balteatus) can be identified as a mature male by the two large rostral horns on his head, ornaments which are used to compete for mates. Endemic to only a small region of Madagascar’s southeastern rainforests, this species has become increasingly endangered and rare due to habitat loss and collection for the pet trade. Ranomafana National Park, Madagascar.
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  • While pythons undoubtedly hold the crown for the largest and most famous constricting snakes of Indonesia, a single genus of boas also occurs in the rainforests of the eastern archipelago. Often overlooked, the Pacific Keel-scaled Boa (Candoia carinata) seldom exceeds a meter in length. It is an ambush predator of frogs and lizards, and like other boas, bears live young rather than laying eggs. Digul River, South Papua, Indonesia (New Guinea).
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  • Borneo Anglehead Lizard (Gonocephalus bornensis). Sarawak, Malaysia (Borneo).
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  • Spotted Cat Snake (Boiga drapiezii). Sarawak, Malaysia (Borneo).
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  • Bornean Angle-headed Lizard (Gonocephalus bornensis), male. Sarawak, Malaysia (Borneo).
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  • Wolf snake (Stegonotus sp.). Highland Papua, Indonesia (New Guinea).
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  • Paradise Tree Snake (Chrysopelea paradisi). Sarawak, Malaysia (Borneo).
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  • The Oriental Vine Snake (Ahaetulla prasina) is an agile climber, feeding on small birds and lizards with its excellent binocular vision. Sabah, Malaysia (Borneo).
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  • Lined Pipe Snake (Cylindrophis lineatus) also known as Blanford's Pipe Snake, a rare species endemic to Borneo and known from only a few specimens. When threatened, this species defends itself by raising its tail which is thickened and colored to appear as a head. Sarawak, Malaysia (Borneo).
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  • The Fire-lipped Keelback (Rhabdophis murudensis) is a rare snake known only from the higher mountains of northern Borneo.  When threatened, it has the remarkable ability to exude a white milky venom from the back of its neck as a defensive mechanism. Gunung Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • Carpet Python (Morelia spilota). Queensland, Australia.
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  • A Pilbara Death Adder (Acanthophis wellsi) from the desert regions of Western Australia. Although superficially resembling vipers by their stout body and habit of ambushing prey, death adders are actually elapids and more closely related to cobras, kraits, and coral snakes. Cape Range National Park, Western Australia.
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  • A master of camouflage, the Mossy Leaf-tailed Gecko (Uroplatus sikorae) is almost completely undetectable when it is at rest on a small tree in the forest understory. Frilled flaps of skin along its body break up its outline, making it appear nearly seamless with the tree on which it rests. Andasibe, Madagascar.
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  • A young Parson's Chameleon (Calumma parsonii) in the rainforest of Masoala National Park. Antsiranana, Madagascar.
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  • Green Tree Python (Morelia azurea), juvenile.
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  • The impressive Sailfin Lizard (Hydrosaurus celebensis) is the largest agamid lizard in the world, reaching over 1 m in length.  They are very agile swimmers and are seldom found far from rivers.  They feed on leaves and insects. Central Sulawesi, Indonesia.
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  • White-lipped Python (Leiopython albertisii). West Papua, Indonesia (New Guinea).
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  • Sap-feeding bugs such as this lanternfly (Pyrops whiteheadi) ingest large quantities of the nutrient-poor fluid, excreting the excess in the form of honeydew. Here a gecko (Gehyra mutilata) waits below the insect for an opportunistic meal of the sweet fluid. Sabah, Malaysia (Borneo).
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  • Southern New Guinea Crocodile (Crocodylus halli), juvenile. Wasur National Park, South Papua, Indonesia (New Guinea).
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  • Three-banded Bridled Snake (Lycodon tristrigatus). Sarawak, Malaysia (Borneo).
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  • Sepik Bent-toed Gecko (Cyrtodactylus novaeguineae). South Papua, Indonesia (New Guinea).
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  • Neotropical Snail-Eater (Dipsas indica). Yasuní National Park, Ecuador.
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  • Smith's Green-eyed Gecko (Gekko smithii). Kinabatangan River, Sabah, Malaysia (Borneo).
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  • Inger's Slug-eating Snake (Asthenodipsas ingeri). Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • Bornean Keeled Pit Viper (Tropidolaemus subannulatus). Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Modest Forest Dragon (Hypsilurus modestus). Papua, Indonesia (New Guinea).
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  • Forest Water Snake (Thamnosophis infrasignatus). Ranomafana National Park, Madagascar.
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  • When threatened, the harmless Ceylonese Pipe Snake (Cylindrophis maculatus) employs a clever trick: it flattens its tail and raises it to expose the bright checkered pattern underneath. Not only does this divert attacks away from its vulnerable head (which remains hidden beneath), but these bold color patterns resemble venomous coral snakes (Calliophis sp.). Central Province, Sri Lanka.
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  • Boulenger's Green Anole (Anolis chloris), male displaying. Choco, Colombia.
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  • A sleeping Short-horned Chameleon (Calumma brevicorne) wakes too slowly to react in time when an unaware spider clambers over its face. Andasibe, Madagascar.
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  • Madagascar Cat-eyed Snake (Madagascarophis colubrinus). Mahajanga, Madagascar.
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  • Peter's Bent-toed Gecko (Cyrtodactylus consobrinus). Sarawak, Malaysia (Borneo).
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