Chien C. Lee

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  • Like other small animals of the forest understory, this Borneo Forest Dragon (Gonocephalus bornensis) must juggle his attention between searching for prey (insects and spiders) while keeping a watchful eye out for larger predators. Although this species is unable to glide like the closely-related Dracos, it is nevertheless an agile climber and can move swiftly up and down trees. The impressive mane on the back of this male signifies his maturity - we will readily defend his territory against rival males. Danum Valley, Sabah, Malaysia (Borneo).
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  • Kuhl's Gliding Gecko (Gekko kuhli). By means of its large webbed feet and lateral skin flaps, this arboreal gecko is capable of gliding or parachuting between trees. Sarawak, Malaysia (Borneo).
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  • Kuhl's Gliding Gecko (Gekko kuhli). Equipped with webbed feet and parasail-like flaps of skin all along its body, this tree-dwelling gecko is able to ‘parachute’ when leaping through the air to escape predators. This not only softens its landing but also enables it to glide a considerable distance. Sarawak, Malaysia (Borneo).
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  • The superbly camouflaged Kinabalu Gliding Gecko (Gekko rhacophorus) is endemic to the montane forests of Mount Kinabalu in northern Borneo.  This poorly-known species is a nocturnal insectivorous tree-dweller and, like other geckos of this genus, possesses skin flaps to assist in a parachuting flight. Sabah, Malaysia (Borneo).
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  • Black-lipped Shrub Lizard (Pelturagonia nigrilabris). Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Horsfield's Gliding Gecko (Gekko horsfieldii). Sabah, Malaysia (Borneo).
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  • A species of bent-toed gecko (Cyrtodactylus cf. mikianus) from the Mamberamo Basin in western New Guinea. Papua, Indonesia.
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  • When threatened, the Knob-tailed Gecko (Nephrurus levis) raises its body off the ground to appear larger in size. Cape Range National Park, Western Australia.
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  • Common Sun Skink (Eutropis multifasciata). Sarawak, Malaysia (Borneo).
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  • The Spiny-headed Shrub Lizard (Pelturagonia spiniceps) is known only from montane forest at a few localities in northwestern Borneo. Pulong Tau National Park, Sarawak, Malaysia (Borneo).
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  • Four-clawed Gecko (Gehyra mutilata) attending a lanternfly (Pyrops whiteheadi) for secreted honeydew. Host tree is Pometia pinnata. Sabah, Malaysia (Borneo).
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  • Madagascar Giant Day Gecko (Phelsuma grandis). Canal des Pangalanes, Madagascar.
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  • Black-lipped Shrub Lizard (Pelturagonia nigrilabris). Sabah, Malaysia (Borneo).
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  • Cryptic Chameleon (Calumma crypticum), male. Ranomafana National Park, Madagascar.
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  • Blue-eyed Angle-headed Lizard (Gonocephalus liogaster), male. Sarawak, Malaysia (Borneo).
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  • A juvenile Parson's Chameleon (Calumma parsoni). More frequently occurring in shades of green, this is one of the world's largest chameleons, with adult specimens sometimes reaching the size of a house cat. They are endemic to rainforest habitats on Madagascar and are threatened by both habitat loss and collection for the international pet trade. Andasibe, Madagascar.
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  • Spiny-tailed Gecko (Diplodactylus ciliaris). Exmouth, Western Australia.
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  • A juvenile Rock Monitor (Varanus albigularis). Mpumalanga, South Africa.
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  • Ornate Earless Agama (Aphaniotis ornata). Sabah, Malaysia (Borneo).
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  • A large male O'Shaughnessy's Chameleon (Calumma oshaughnessyi) stalks up a forest liana, keeping an eye out for both prey and predator alike. With 360-degree vision, remarkable image acuity, and monocular depth perception, chameleons can detect insects that might be well out of reach, judge their distance accurately, and even distinguishing them from their surroundings when the insect is completely still. Ranomafana National Park, Madagascar.
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  • Purple Anole (Anolis purpurescens). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • Horned Flying Lizard (Draco cornutus), male using his dewlap to display for courtship and territorial aggression. Sabah, Malaysia (Borneo).
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  • A young Parson's Chameleon (Calumma parsonii) in the rainforest of Masoala National Park. Antsiranana, Madagascar.
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  • Carefully sequestered on a perch not far above the rainforest floor, a Moyer’s Pygmy Chameleon (Rhampholeon moyeri) sleeps by night safely hidden from prowling nocturnal predators. Measuring less than 6 cm in total length, these are some of the smallest and most camouflaged in Africa – by day when they hunt for insects on the ground they are nearly invisible among the leaf litter. Recent research has shown that there is a surprising amount of diversity among Rhampholeon chameleons (26 currently recognized species), many of which are restricted to their own isolated mountain ranges. . Udzungwa Mountains, Tanzania.
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  • Sumatran Flying Dragon (Draco sumatranus), male performing territorial display. Sarawak, Malaysia (Borneo).
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  • The mimicry of a dead leaf is so precise for the Satanic Leaf-tailed Gecko (Uroplatus phantasticus), they can often hide effectively in plain sight. It has been untested to what degree these geckos actively select a substrate that matches their individual patterns, but U. phantasticus appears to typically hide among dead foliage which suits it perfectly. Ranomafana National Park, Madagascar.
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  • Long-tailed Grass Lizard (Takydromus sexlineatus). Sarawak, Malaysia (Borneo).
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  • The edges of this leaf-tailed gecko (Uroplatus sikorae) seem to blend seamlessly with the branch on which it rests. Ranomafana National Park, Madagascar.
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  • A sleeping Short-horned Chameleon (Calumma brevicorne) wakes too slowly to react in time when an unaware spider clambers over its face. Andasibe, Madagascar.
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  • The Bornean Forest Dragon (Gonocephalus bornensis) is a large arboreal agamid lizard endemic to the rainforests of Borneo. It feeds on ants and spiders. Sarawak, Malaysia (Borneo).
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  • The Earless Monitor Lizard (Lanthanotus borneensis) is endemic to Borneo where it lives in rocky stream habitats in lowland rainforest. Because of its extreme rarity (it is known from only a few specimens), its life history remains unstudied. It is the only species within its entire family (Lanthanotidae) but it is related to the true monitor lizards (Varanidae). Sarawak, Malaysia (Borneo).
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  • Closeup portrait of a Tokay Gecko (Gekko gecko). Uthai Thani, Thailand.
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  • A study on perfection in mimicry: a Satanic Leaf-tailed Gecko (Uroplatus phantasticus). One of 14 species of leaf-tailed geckos endemic to Madagascar, this nocturnal insectivore utilizes its incredible camouflage to hide among dead foliage by day. All Uroplatus geckos are under severe threat from deforestation and over-collecting for the international pet trade. Andasibe, Madagascar.
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  • Draco cornutus, in flight. Flying lizards of the genus Draco are superbly adapted for their arboreal lifestyle. They are capable of gliding from tree to tree on wing-like flaps of skin supported by their movable ribs. They feed primarily on ants. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • With colors that seem to glow with an unearthly radiance, a male Osa Anole (Anolis osa) betrays his normally cryptic appearance to briefly flash his extended dewlap. Anole lizards are famous for these flamboyant visual signals – they are used to attract mates, ward off rivals, and even sometimes dissuade potential predators. A 2015 study found that the translucent quality of anole dewlaps was a crucial component in their ability to transmit these color signals effectively, particularly in shady environments; under the right conditions they possess an almost unnatural luminance. This species, which is endemic to Costa Rica’s Osa Peninsula, spends most of its time on perches near ground level in the dappled light of the rainforest understory. Osa Peninsula, Puntarenas, Costa Rica.
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  • With night vision hundreds of times more sensitive than ours and the ability to discern color even in near total darkness, Madagascar’s leaf-tailed geckos (Uroplatus spp.) are superbly adapted as nocturnal insect hunters. Large eyes, however, demand special maintenance. Like most geckos, Uroplatus have no eyelids and cannot blink; instead, they utilize their long tongues like windshield wipers, keeping their eyes moist and clean. This is a young male Satanic Leaf-tailed Gecko (U. phantasticus), a denizen of the island’s rainforest zone and a superb mimic of dead foliage. Ranomafana National Park, Madagascar.
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  • The mimicry of a dead leaf is so precise for the Satanic Leaf-tailed Gecko (Uroplatus phantasticus), they can often hide effectively in plain sight. It has been untested to what degree these geckos actively select a substrate that matches their individual patterns, but U. phantasticus appears to typically hide among dead foliage which suits it perfectly. Ranomafana National Park, Madagascar.
    cld1916615.jpg
  • Males of the Madagascar's Satanic Leaf-tailed Gecko (Uroplatus phantasticus) can usually be distinguished by their elaborately notched tails, a trait which gives them better mimicry of a leaf that has been partially eaten by insects. Exactly why this is a sexually dimorphic feature in these geckos is unknown, but in some animals different camouflage patterns in males and females can arise where each sex spends their time hiding in different locations and thus face differing selective pressures on their appearance. Ranomafana National Park, Madagascar.
    cld1917741.jpg
  • Few creatures possess a mastery of camouflage the way Madagascar’s leaf-tailed geckos do. This Mossy Leaf-tailed Gecko (Uroplatus sikorae) sleeps by day on a rainforest liana, its outline obscured by frilled dermal flaps that line its body. By always resting in a downward-facing position they may be able to quickly catch unsuspecting insects that climb up from the ground and also conceal their reflective lidless eyes from predators lurking above. Ranomafana National Park, Madagascar.
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  • Although masterful visual predators by day, chameleons lack light-sensitive rods in their eyes and thus have poor night vision. Because of this they typically retire at dusk to a sleeping perch, often returning to a favorite exposed branch which ideally puts them out of the reach of nocturnal snakes. This dozing Two-banded Chameleon (Furcifer balteatus) can be identified as a mature male by the two large rostral horns on his head, ornaments which are used to compete for mates. Endemic to only a small region of Madagascar’s southeastern rainforests, this species has become increasingly endangered and rare due to habitat loss and collection for the pet trade. Ranomafana National Park, Madagascar.
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  • Komodo Dragons (Varanus komodoensis), like all monitor lizards, posses forked tongues which are used to smell the air (chemoreception) in a similar fashion to snakes. Rinca Island, East Nusa Tenggara, Indonesia.
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  • Two-banded Chameleon (Furcifer balteatus), female. Ranomafana National Park, Madagascar.
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  • An adult Common Butterfly Lizard (Leiolepis belliana) keeps a watchful eye out for predators whilst her young remain at the entrance of the nest burrow.  Nine species of butterfly lizards are known from the drier sandy regions of Southeast Asia, but their life-history is poorly documented. Uthai Thani, Thailand.
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  • Northern Leaf-tailed Gecko (Saltuarius cornutus). Queensland, Australia.
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  • Hikida's Forest Dragon (Hypsilurus hikidanus). Central Papua, Indonesia (New Guinea).
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  • Sap-feeding bugs such as this lanternfly (Pyrops whiteheadi) ingest large quantities of the nutrient-poor fluid, excreting the excess in the form of honeydew. Here a gecko (Gehyra mutilata) waits below the insect for an opportunistic meal of the sweet fluid. Sabah, Malaysia (Borneo).
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  • Sepik Bent-toed Gecko (Cyrtodactylus novaeguineae). South Papua, Indonesia (New Guinea).
    cld1716408.jpg
  • Large-headed Shrub Lizard (Pelturagonia cephalum). Kinabalu National Park, Sabah, Malaysia (Borneo).
    cld1800488.jpg
  • Modest Forest Dragon (Hypsilurus modestus). Papua, Indonesia (New Guinea).
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  • Mossy Leaf-tailed Gecko (Uroplatus sikorae). Mantadia National Park, Madagascar.
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  • Boulenger's Green Anole (Anolis chloris), male displaying. Choco, Colombia.
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  • Two-banded Chameleon (Furcifer balteatus), female. Ranomafana National Park, Madagascar.
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  • Peter's Bent-toed Gecko (Cyrtodactylus consobrinus). Sarawak, Malaysia (Borneo).
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  • Doria's Angle-headed Lizard (Gonocephalus doriae). Sarawak, Malaysia (Borneo).
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  • The rainforests of the remote Marojejy National Park in northern Madagascar are one of the best places to look for the island's unique leaf-tailed geckos. In addition to their superb camouflage, most species, like this Uroplatus giganteus, rest head downwards presumably to conceal the revealing reflections of their lidless eyes from arboreal predators.
    cld1621209.jpg
  • Smith's Green-eyed Gecko (Gekko smithii). Kinabatangan River, Sabah, Malaysia (Borneo).
    cld10010450.jpg
  • With over one hundred species occurring throughout the island in virtually all habitats, Madagascar is a hotspot for gecko diversity and endemism. This Graceful Madagascar Ground Gecko (Paroedura gracilis) is an inhabitant of the eastern rainforests and, unlike most arboreal species, lacks the enlarged toe pads for climbing vertical surfaces. Active only at night, it hunts insects from low perches in the forest understory. Marojejy National Park, Madagascar.
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  • A Lined Gecko (Gekko vittatus) perches in its rainforest habitat on the island of Halmahera, central Indonesia. Sometimes referred to as the "Skunk Gecko" because of its distinctive coloration, these are popular animals in the pet trade and large numbers are collected and exported yearly from Indonesia and the Solomon Islands. North Maluku, Indonesia.
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  • Sharp-nosed Chameleon (Kinyongia oxyrhina), male. Udzungwa Mountains, Tanzania.
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  • Round-eyed Gecko (Cnemaspis sp. aff. paripari), undescribed and one of several species of Cnemaspis which are endemic to separate and isolated limestone outrops. Sarawak, Malaysia (Borneo).
    cld2102784.jpg
  • Tug of war: a Giant Huntsman Spider (Heteropoda sp.) struggles to keep a hold on her precious egg sac as a hungry Banded Forest Gecko (Cyrtodactylus consobrinus) attempts to steal it away. With sometimes over 200 eggs within the papery sac, the female spider guards her eggs ferociously, carrying the bundle in her jaws for several weeks until they hatch. Fortunately for the gecko, this spider seems more concerned with maintaining her grip with her mandibles rather than risking a quick release to bite the offending gecko. Sarawak, Malaysia (Borneo).
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  • Broad-headed Woodlizard (Enyalioides laticeps). Yasuní National Park, Ecuador.
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  • Huntsman spider (Sparassidae) preying on a young Madagascar day gecko (Phelsuma parva). Tomasina, Madagascar.
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  • Kirindy Leaf-toed Gecko (Paroedura rennerae), juvenile. Tsingy de Bemaraha National Park, Madagascar.
    cld1720970.jpg
  • The Earless Monitor Lizard (Lanthanotus borneensis) is endemic to Borneo where it lives in rocky stream habitats in lowland rainforest. Because of its extreme rarity (it is known from only a few specimens), its life history remains unstudied. It is the only species within its entire family (Lanthanotidae) and was once though to be the missing link between lizards and snakes. Sarawak, Malaysia (Borneo).
    cld1704529.jpg
  • Horned Gliding Lizard (Draco cornutus), male displaying gular flap. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
    cld1401802.jpg
  • Satanic Leaf-tailed Gecko (Uroplatus phantasticus), female. Ranomafana National Park, Madagascar.
    cld1622883.jpg
  • Males of the Madagascar's Satanic Leaf-tailed Gecko (Uroplatus phantasticus) can usually be distinguished by their elaborately notched tails, a trait which gives them better mimicry of a leaf that has been partially eaten by insects. Exactly why this is a sexually dimorphic feature in these geckos is unknown, but in some animals different camouflage patterns in males and females can arise where each sex spends their time hiding in different locations and thus face differing selective pressures on their appearance. Ranomafana National Park, Madagascar.
    cld1917721.jpg
  • Borneo Anglehead Lizard (Gonocephalus bornensis). Sarawak, Malaysia (Borneo).
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  • Bornean Angle-headed Lizard (Gonocephalus bornensis), male. Sarawak, Malaysia (Borneo).
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  • Short-horned Chameleon (Calumma brevicorne), female. Andasibe, Madagascar.
    cld1507473.jpg
  • A master of camouflage, the Mossy Leaf-tailed Gecko (Uroplatus sikorae) is almost completely undetectable when it is at rest on a small tree in the forest understory. Frilled flaps of skin along its body break up its outline, making it appear nearly seamless with the tree on which it rests. Andasibe, Madagascar.
    cld1508652.jpg
  • The impressive Sailfin Lizard (Hydrosaurus celebensis) is the largest agamid lizard in the world, reaching over 1 m in length.  They are very agile swimmers and are seldom found far from rivers.  They feed on leaves and insects. Central Sulawesi, Indonesia.
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  • Natalia's Spiny Lizard (Acanthosaura nataliae). Many lizards sport spines for display or protection, but Acanthosaura take this development to an extreme. Despite their fearsome appearance these arboreal reptiles still fall prey to certain snakes, and thus need to sleep on thin branches such as this during the night. Thua Thien Hue, Vietnam.
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  • Satanic Leaf-tailed Gecko (Uroplatus phantasticus). Ranomafana National Park, Madagascar.
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  • Cat Gecko (Aeluroscalabotes felinus). These slow-moving and normally extremely placid reptiles only put on an aggressive display when their other defenses (camouflage and tail dropping) have failed them. Members of the ‘eyelid geckos’, they are the only representative of the family Eublepharidae in Borneo. Sarawak, Malaysia.
    cld1701081.jpg
  • A master of camouflage, the Mossy Leaf-tailed Gecko (Uroplatus sikorae) is almost completely undetectable when it is at rest on a small tree in the forest understory. Frilled flaps of skin along its body break up its outline, making it appear nearly seamless with the tree on which it rests. Andasibe, Madagascar.
    cld1508663.jpg
  • Rough Chameleon (Trioceros rudis). Ruhengeri, Rwanda.
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  • Dusky Gliding Lizard (Draco obscurus). Sarawak, Malaysia (Borneo).
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  • Satanic Leaf-tailed Gecko (Uroplatus phantasticus). Andasibe, Madagascar.
    cld1716737.jpg
  • The rainforests of the remote Marojejy National Park in northern Madagascar are one of the best places to look for the island's unique leaf-tailed geckos. In addition to their superb camouflage, most species, like this Uroplatus giganteus, rest head downwards presumably to conceal the revealing reflections of their lidless eyes from arboreal predators.
    cld1621379.jpg
  • Dumeril's Monitor (Varanus dumerilii), juvenile. Sarawak, Malaysia (Borneo).
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  • Spiny Ground Gecko (Paroedura bastardi). Amboasary, Madagascar.
    cld1614839.jpg
  • This female Madagascar Pygmy Kingfisher (Corythornis madagascariensis) has just been gifted a small lizard (Madascincus melanopleura) by her mate as part of a courtship ritual. Despite their name, these birds are not dependent on water, and typically hunt for insects and small vertebrates on the forest floor. Ranomafana National Park, Madagascar.
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  • Boulenger's Tree Agama (Dendragama boulengeri). Kerinci Seblat National Park, West Sumatra, Indonesia.
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  • Greater Carpet Chameleon (Furcifer major). Isalo National Park, Madagascar.
    cld1832594.jpg
  • The Earless Monitor Lizard (Lanthanotus borneensis) is endemic to Borneo where it lives in rocky stream habitats in lowland rainforest. Because of its extreme rarity (it is known from only a few specimens), its life history remains unstudied. It is the only species within its entire family (Lanthanotidae) but it is related to the true monitor lizards (Varanidae). Sarawak, Malaysia (Borneo).
    cld1308209.jpg
  • Graceful Madagascar Ground Gecko (Paroedura gracilis). Marojejy National Park, Madagascar.
    cld1621946.jpg
  • Leaf Anole (Anolis planiceps), male. Bolivar, Venezuela.
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  • Giant Leaf-tailed Gecko (Uroplatus fimbriatus) from Masoala National Park. Antsiranana, Madagascar.
    cld1509427.jpg
  • Two-banded Chameleon (Furcifer balteatus), male. A rare species, F. balteatus is known with certainty to occur at only a few locations in southeast Madagascar, and is classified as Endangered due to threats from habitat loss and collecting for the pet trade. Ranomafana National Park, Madagascar.
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  • Borneo Forest Dragon (Gonocephalus bornensis). Sabah, Malaysia (Borneo).
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  • Whistling Lizard (Calotes liolepis). Central Province, Sri Lanka.
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  • Warty Chameleon (Furcifer verrucosus). Toliara, Madagascar.
    cld1833118.jpg
  • Undoubtedly one of the rarest and most peculiar of all Borneo's reptiles: the Bornean Horned Lizard (Harpesaurus borneensis). The males of this species are unique in possessing a distinctive horn at the tip of their snout, a trait not seen in the females nor found in any other Bornean lizard. Even more unusual, this is the only known agamid lizard in Southeast Asia to give birth to live young rather than laying eggs. As only a few specimens of this species have ever been found, its full life history and distribution in Borneo are still largely unknown. Sarawak, Malaysia (Borneo).
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  • Desert Iguana (Dipsosaurus dorsalis). Baja California Norte, Mexico.
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  • Being as discreet as possible, a tiny Elongate Leaf Chameleon (Palleon nasus) makes its way across the forest floor at a snail’s pace, freezing whenever it feels someone watching. The adults of this species measure less than 9cm long and are at the small end of Madagascar’s chameleon spectrum. Their camouflage is a perfect adaptation for their specific habitat: the leaf litter of the rainforest floor. Ranomafana National Park, Madagascar.
    cld1837043.jpg
  • Endemic to the mountains of central Sri Lanka, the Rhino Horn Lizard (Ceratophora stoddartii) is named for the unusual spine found on the snout of males. This is likely a product of female mate selection. Central Province, Sri Lanka.
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  • Blotch Bow-finger Gecko (Cyrtodactylus yakhuna). North Western Province, Sri Lanka.
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  • Hump-nosed lizard photo (Lyriocephalus scutatus), male. Sinharaja National Park, Southern Province, Sri Lanka.
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