Chien C. Lee

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  • This small dung beetle (Paragymnopleurus maurus) is preparing a ball made from monkey droppings in which it will deposit its eggs and then bury. Sarawak, Malaysia (Borneo).
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  • Dung beetle (Paragymnopleurus maurus), rolling scat of Sambar Deer. Danum Valley, Sabah, Malaysia (Borneo).
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  • Dung beetle (Paragymnopleurus sparsus), rolling scat of Sambar Deer. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Dung Beetle (Epilissus splendidus). Mantadia National Park, Madagascar.
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  • Dung beetle (Scarabaeus sp.), rolling dung. Bundala National Park, Sri Lanka.
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  • A pair of the largest dung beetles in Borneo (Catharsius dayacus) clean up after the droppings of a Sambar Deer in the rainforest of Danum Valley, Sabah. Although this species does not roll balls like its more well-known relatives, it nevertheless is responsible for burying large amounts of animal scat and provides a vital ecological service. Nearly a hundred species of dung beetles have been recorded from Danum alone; studies have shown that their diversity not only directly reflects the wide assortment of vertebrates in an area but also can serve as a valuable indicator of habitat heterogeneity and forest disturbance. Sabah, Malaysia (Borneo).
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  • Dung beetle (Proagoderus watanabei), male feeding on herbivore scat. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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