Chien C. Lee

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  • Flat-nosed Pitviper (Craspedocephalus puniceus). West Java, Indonesia.
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  • Black-lipped Shrub Lizard (Pelturagonia nigrilabris). Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Kuhl's Gliding Gecko (Gekko kuhli). By means of its large webbed feet and lateral skin flaps, this arboreal gecko is capable of gliding or parachuting between trees. Sarawak, Malaysia (Borneo).
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  • Kuhl's Gliding Gecko (Gekko kuhli). Equipped with webbed feet and parasail-like flaps of skin all along its body, this tree-dwelling gecko is able to ‘parachute’ when leaping through the air to escape predators. This not only softens its landing but also enables it to glide a considerable distance. Sarawak, Malaysia (Borneo).
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  • Triangle Keelback (Xenochrophis trianguligerus). Maliau Basin Conservation Area, Sabah, Malaysia (Borneo).
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  • The rare Kinabalu Brown Pit Viper (Garthius chaseni) is known only from a few mountains in northern Borneo where it occurs in high elevation forest. Very little is known of its life history as it has seldom been observed in the wild. Sabah, Malaysia (Borneo).
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  • Still wet from a passing rainstorm, a Sabah Bamboo Pitviper (Trimeresurus sabahi) lies in ambush on the side of a tree, waiting for the approach of suitable prey. By maintaining a low metabolism and sometimes remaining in a single position for weeks at a time, tropical vipers such as this can survive by feeding as infrequently as once every few months. Mount Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • Black-lipped Shrub Lizard (Pelturagonia nigrilabris). Sabah, Malaysia (Borneo).
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  • Large-headed Shrub Lizard (Pelturagonia cephalum). Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • The superbly camouflaged Kinabalu Gliding Gecko (Gekko rhacophorus) is endemic to the montane forests of Mount Kinabalu in northern Borneo.  This poorly-known species is a nocturnal insectivorous tree-dweller and, like other geckos of this genus, possesses skin flaps to assist in a parachuting flight. Sabah, Malaysia (Borneo).
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  • Sabah Bamboo Pitviper (Trimeresurus sabahi). Sarawak, Malaysia (Borneo).
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  • Flat-nosed Pitviper (Craspedocephalus puniceus). West Java, Indonesia.
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  • When threatened, the Knob-tailed Gecko (Nephrurus levis) raises its body off the ground to appear larger in size. Cape Range National Park, Western Australia.
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  • White-spotted Cat Snake (Boiga drapiezii), green morph. Sarawak, Malaysia (Borneo).
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  • Bornean Short Python (Python breitensteini). Sarawak, Malaysia (Borneo).
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  • The mimicry of a dead leaf is so precise for the Satanic Leaf-tailed Gecko (Uroplatus phantasticus), they can often hide effectively in plain sight. It has been untested to what degree these geckos actively select a substrate that matches their individual patterns, but U. phantasticus appears to typically hide among dead foliage which suits it perfectly. Ranomafana National Park, Madagascar.
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  • Purple Anole (Anolis purpurescens). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • Malayan Banded Wolf Snake (Lycodon subcinctus), juvenile. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Long-tailed Grass Lizard (Takydromus sexlineatus). Sarawak, Malaysia (Borneo).
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  • Brown-tailed Racer (Gonyosoma oxycephalum). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Like other small animals of the forest understory, this Borneo Forest Dragon (Gonocephalus bornensis) must juggle his attention between searching for prey (insects and spiders) while keeping a watchful eye out for larger predators. Although this species is unable to glide like the closely-related Dracos, it is nevertheless an agile climber and can move swiftly up and down trees. The impressive mane on the back of this male signifies his maturity - we will readily defend his territory against rival males. Danum Valley, Sabah, Malaysia (Borneo).
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  • Few creatures possess a mastery of camouflage the way Madagascar’s leaf-tailed geckos do. This Mossy Leaf-tailed Gecko (Uroplatus sikorae) sleeps by day on a rainforest liana, its outline obscured by frilled dermal flaps that line its body. By always resting in a downward-facing position they may be able to quickly catch unsuspecting insects that climb up from the ground and also conceal their reflective lidless eyes from predators lurking above. Ranomafana National Park, Madagascar.
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  • Madagascar Giant Day Gecko (Phelsuma grandis). Canal des Pangalanes, Madagascar.
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  • Endemic to the mountains of central Sri Lanka, the Rhino Horn Lizard (Ceratophora stoddartii) is named for the unusual spine found on the snout of males. This is likely a product of female mate selection. Central Province, Sri Lanka.
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  • Horned Flying Lizard (Draco cornutus), male using his dewlap to display for courtship and territorial aggression. Sabah, Malaysia (Borneo).
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  • Undoubtedly one of the rarest and most peculiar of all Borneo's reptiles: the Bornean Horned Lizard (Harpesaurus borneensis). The males of this species are unique in possessing a distinctive horn at the tip of their snout, a trait not seen in the females nor found in any other Bornean lizard. Even more unusual, this is the only known agamid lizard in Southeast Asia to give birth to live young rather than laying eggs. As only a few specimens of this species have ever been found, its full life history and distribution in Borneo are still largely unknown. Sarawak, Malaysia (Borneo).
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  • Cryptic Chameleon (Calumma crypticum), male. Ranomafana National Park, Madagascar.
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  • Ecuadorian Toadheaded Pitviper (Bothrocophias campbelli). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • Mocquard's Madagascar Ground Gecko (Paroedura bastardi), juvenile. Tsingy de Bemaraha National Park, Madagascar.
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  • The Twin-barred Flying Snake (Chrysopelea pelias) is the smallest and rarest of Southeast Asia's flying snake species. Like other Chrysopelea, this snake hunts lizards and is almost completely arboreal. Sarawak, Malaysia (Borneo).
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  • The Earless Monitor Lizard (Lanthanotus borneensis) is endemic to Borneo where it lives in rocky stream habitats in lowland rainforest. Because of its extreme rarity (it is known from only a few specimens), its life history remains unstudied. It is the only species within its entire family (Lanthanotidae) and was once though to be the missing link between lizards and snakes. Sarawak, Malaysia (Borneo).
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  • Although the Banded Coral Snake (Calliophis intestinalis) is not agressive, it possesses an extremely toxic venom like other Elapids. When disturbed it displays its red tail and the brightly colored bands on its undersurface which serve as a warning signal to would-be predators. Sarawak, Malaysia (Borneo).
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  • Paradise Tree Snake (Chrysopelea paradisi). Sarawak, Malaysia (Borneo).
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  • Blue-eyed Angle-headed Lizard (Gonocephalus liogaster), male. Sarawak, Malaysia (Borneo).
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  • Paradise Tree Snake (Chrysopelea paradisi). Sarawak, Malaysia (Borneo).
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  • Lined Pipe Snake (Cylindrophis lineatus) also known as Blanford's Pipe Snake, a rare species endemic to Borneo and known from only a few specimens. When threatened, this species defends itself by raising its tail which is thickened and colored to appear as a head. Sarawak, Malaysia (Borneo).
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  • A juvenile Parson's Chameleon (Calumma parsoni). More frequently occurring in shades of green, this is one of the world's largest chameleons, with adult specimens sometimes reaching the size of a house cat. They are endemic to rainforest habitats on Madagascar and are threatened by both habitat loss and collection for the international pet trade. Andasibe, Madagascar.
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  • Bronzeback (Dendrelaphis terrificus). Gorontalo, Sulawesi, Indonesia.
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  • Spiny-tailed Gecko (Diplodactylus ciliaris). Exmouth, Western Australia.
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  • A study on perfection in mimicry: a Satanic Leaf-tailed Gecko (Uroplatus phantasticus). One of 14 species of leaf-tailed geckos endemic to Madagascar, this nocturnal insectivore utilizes its incredible camouflage to hide among dead foliage by day. All Uroplatus geckos are under severe threat from deforestation and over-collecting for the international pet trade. Andasibe, Madagascar.
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  • The Bornean Keeled Green Pit Viper (Tropidolaemus subannulatus) is an arboreal species, often laying in wait for prey on the same branch for weeks at a time. Sarawak, Malaysia (Borneo).
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  • Hikida's Forest Dragon (Hypsilurus hikidanus). Papua, Indonesia (New Guinea).
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  • Black-headed Cat Snake (Boiga nigriceps). Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Although usually considered a terrestrial species, the Striped Kukri Snake (Oligodon octolineatus) is an excellent climber and can occasionally be seen climbing tree trunks. Sarawak, Malaysia (Borneo).
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  • Sumatran Pit Viper (Trimeresurus sumatranus). Sarawak, Malaysia (Borneo).
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  • Hump-nosed lizard photo (Lyriocephalus scutatus), male. Sinharaja National Park, Southern Province, Sri Lanka.
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  • While pythons undoubtedly hold the crown for the largest and most famous constricting snakes of Indonesia, a single genus of boas also occurs in the rainforests of the eastern archipelago. Often overlooked, the Pacific Keel-scaled Boa (Candoia carinata) seldom exceeds a meter in length. It is an ambush predator of frogs and lizards, and like other boas, bears live young rather than laying eggs. Digul River, Papua, Indonesia (New Guinea).
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  • Borneo Anglehead Lizard (Gonocephalus bornensis). Sarawak, Malaysia (Borneo).
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  • Cloudy Snail-eating Snake (Sibon nebulatus). Limón, Costa Rica.
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  • Common Sun Skink (Eutropis multifasciata). Sarawak, Malaysia (Borneo).
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  • Spiny Ground Gecko (Paroedura bastardi). Amboasary, Madagascar.
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  • This female Madagascar Pygmy Kingfisher (Corythornis madagascariensis) has just been gifted a small lizard (Madascincus melanopleura) by her mate as part of a courtship ritual. Despite their name, these birds are not dependent on water, and typically hunt for insects and small vertebrates on the forest floor. Ranomafana National Park, Madagascar.
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  • Catesby's Snail-eater (Dipsas catesbyi). Yasuní National Park, Ecuador.
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  • Red-tailed Pipe Snake (Cylindrophis ruffus). Sarawak, Malaysia (Borneo).
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  • Four-clawed Gecko (Gehyra mutilata) attending a lanternfly (Pyrops whiteheadi) for secreted honeydew. Host tree is Pometia pinnata. Sabah, Malaysia (Borneo).
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  • Amazon Tree Boa (Corallus hortulanus). Yasuní National Park, Ecuador.
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  • Scarcely longer than an earthworm, this juvenile slug-eating snake (Pareas hamptoni) sleeps balanced on top of a fern frond. Many slug-eating snakes retain the  unusual asymmetric dentition of their snail-eating ancestors, an adaptation for pulling snails out of their shells via a process known as ‘mandibular walking’. Bach Ma National Park, Vietnam.
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  • Vogel’s Pit Viper (Trimeresurus vogeli). Bach Ma National Park, Vietnam.
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  • The Three-striped Ground Snake (Pseudoxyrhopus tritaeniatus) is certainly one of the most beautiful of Madagascan endemic snakes, but why is it colored this way? Bold red stripes are frequently used as an aposematic warning, indicating to other animals of severe danger. It is curious then that not only is this species completely non-venomous but there are in fact no other dangerous snakes in Madagascar that this could possibly be a mimic of. An interesting theory points to the co-occurrence of large red toxic millipedes in the same forests – could these actually be a mimicry model for this snake? On an island legendary for its biological curiosities, nothing seems beyond the realm of possibility. Ranomafana National Park, Madagascar.
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  • Males of the Madagascar's Satanic Leaf-tailed Gecko (Uroplatus phantasticus) can usually be distinguished by their elaborately notched tails, a trait which gives them better mimicry of a leaf that has been partially eaten by insects. Exactly why this is a sexually dimorphic feature in these geckos is unknown, but in some animals different camouflage patterns in males and females can arise where each sex spends their time hiding in different locations and thus face differing selective pressures on their appearance. Ranomafana National Park, Madagascar.
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  • Mossy Leaf-tailed Gecko (Uroplatus sikorae). Mantadia National Park, Madagascar.
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  • Cloudy Snail-eating Snake (Sibon nebulatus). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • Blotch Bow-finger Gecko (Cyrtodactylus yakhuna). North Western Province, Sri Lanka.
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  • Whistling Lizard (Calotes liolepis). Central Province, Sri Lanka.
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  • Barnes' Cat Snake (Boiga barnesii). Sinharaja National Park, Sri Lanka.
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  • Hump-nosed Viper (Hypnale zara). Sinharaja National Park, Sri Lanka.
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  • When threatened, the harmless Ceylonese Pipe Snake (Cylindrophis maculatus) employs a clever trick: it flattens its tail and raises it to expose the bright checkered pattern underneath. Not only does this divert attacks away from its vulnerable head (which remains hidden beneath), but these bold color patterns resemble venomous coral snakes (Calliophis sp.). Central Province, Sri Lanka.
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  • Brown Tree Snake (Boiga irregularis), juvenile. Mamberamo Basin, Papua, Indonesia (New Guinea).
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  • Red-headed Reed Snake (Calamaria schlegeli schlegeli), a small soil-dwelling species which feeds on frogs and slugs. Sarawak, Malaysia (Borneo).
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  • Blunthead Tree Snake (Imantodes cenchoa), a lizard hunting specialist that due to its extremely slender body is able to seek out sleeping lizards on even the most delicate of branches. Yasuní National Park, Ecuador.
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  • The bold coloration of the Red-headed Krait (Bungarus flaviceps) is a warning to would-be predators of the highly potent venom it possesses. This large elapid feeds primarily on other reptiles including lizards and snakes. Sarawak, Malaysia (Borneo).
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  • Although masterful visual predators by day, chameleons lack light-sensitive rods in their eyes and thus have poor night vision. Because of this they typically retire at dusk to a sleeping perch, often returning to a favorite exposed branch which ideally puts them out of the reach of nocturnal snakes. This dozing Two-banded Chameleon (Furcifer balteatus) can be identified as a mature male by the two large rostral horns on his head, ornaments which are used to compete for mates. Endemic to only a small region of Madagascar’s southeastern rainforests, this species has become increasingly endangered and rare due to habitat loss and collection for the pet trade. Ranomafana National Park, Madagascar.
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  • Although completely non-venomous, the Painted Mock Viper (Psammodynastes pictus) defends itself by aggressively striking if threatened. It often hunts aquatic animals such as frogs and fish by perching on a low branch over the surface of the water. Sarawak, Malaysia (Borneo).
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  • Spotted Cat Snake (Boiga drapiezii). Sarawak, Malaysia (Borneo).
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  • A Harlequin Gliding Frog (Rhacophorus pardalis) struggles futilely as it is being slowly swallowed alive by a Black-Headed Cat Snake (Boiga nigriceps). Stagnant pools of water in the Borneo rainforest serve as important breeding grounds for many tree frogs which gather around in great numbers, a fact that some predators wisely take advantage of. Kubah National Park, Sarawak, Malaysia (Borneo).
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  • Bornean Angle-headed Lizard (Gonocephalus bornensis), male. Sarawak, Malaysia (Borneo).
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  • Horsfield's Gliding Gecko (Gekko horsfieldii). Sabah, Malaysia (Borneo).
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  • Wolf snake (Stegonotus sp.). Papua, Indonesia (New Guinea).
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  • The Oriental Vine Snake (Ahaetulla prasina) is an agile climber, feeding on small birds and lizards with its excellent binocular vision. Sabah, Malaysia (Borneo).
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  • Common Big-eyed Snake (Mimophis mahfalensis). Toliara, Madagascar.
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  • A juvenile Bornean Dark-necked Slug Snake (Asthenodipsas borneensis). This nocturnal species feeds on slugs and snails in low vegetation. Recently split from A. malaccanus of the Malay Peninsula and Sumatra. Kubah National Park, Sarawak, Malaysia (Borneo).
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  • Two-banded Chameleon (Furcifer balteatus), female. Ranomafana National Park, Madagascar.
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  • The Fire-lipped Keelback (Rhabdophis murudensis) is a rare snake known only from the higher mountains of northern Borneo.  When threatened, it has the remarkable ability to exude a white milky venom from the back of its neck as a defensive mechanism. Gunung Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • Bornean Keeled Green Pit Viper (Tropidolaemus subannulatus). Sarawak, Malaysia (Borneo).
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  • Ornate Earless Agama (Aphaniotis ornata). Sabah, Malaysia (Borneo).
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  • Carpet Python (Morelia spilota). Queensland, Australia.
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  • A Pilbara Death Adder (Acanthophis wellsi) from the desert regions of Western Australia. Although superficially resembling vipers by their stout body and habit of ambushing prey, death adders are actually elapids and more closely related to cobras, kraits, and coral snakes. Cape Range National Park, Western Australia.
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  • A species of bent-toed gecko (Cyrtodactylus cf. mikianus) from the Mamberamo Basin in western New Guinea. Papua, Indonesia.
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  • A master of camouflage, the Mossy Leaf-tailed Gecko (Uroplatus sikorae) is almost completely undetectable when it is at rest on a small tree in the forest understory. Frilled flaps of skin along its body break up its outline, making it appear nearly seamless with the tree on which it rests. Andasibe, Madagascar.
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  • A young Parson's Chameleon (Calumma parsonii) in the rainforest of Masoala National Park. Antsiranana, Madagascar.
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  • A juvenile Oriental Vine Snake (Ahaetulla prasina) basks in a beam of sunlight in the rainforest of Tangkoko National Park. These highly arboreal snakes are excellent climbers and are adept at hunting lizards, their favored prey. North Sulawesi, Indonesia.
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  • A juvenile Rock Monitor (Varanus albigularis). Mpumalanga, South Africa.
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  • Green Tree Python (Morelia azurea), juvenile.
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  • The impressive Sailfin Lizard (Hydrosaurus celebensis) is the largest agamid lizard in the world, reaching over 1 m in length.  They are very agile swimmers and are seldom found far from rivers.  They feed on leaves and insects. Central Sulawesi, Indonesia.
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  • Banded Malaysian Coralsnake (Calliophis intestinalis). Sarawak, Malaysia (Borneo).
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  • Three-banded Bridled Snake (Lycodon tristrigatus). Sarawak, Malaysia (Borneo).
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  • Bornean Keeled Pit Viper (Tropidolaemus subannulatus). Mulu National Park, Sarawak, Malaysia (Borneo).
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  • A sleeping Short-horned Chameleon (Calumma brevicorne) wakes too slowly to react in time when an unaware spider clambers over its face. Andasibe, Madagascar.
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  • The Bornean Forest Dragon (Gonocephalus bornensis) is a large arboreal agamid lizard endemic to the rainforests of Borneo. It feeds on ants and spiders. Sarawak, Malaysia (Borneo).
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  • Natalia's Spiny Lizard (Acanthosaura nataliae). Many lizards sport spines for display or protection, but Acanthosaura take this development to an extreme. Despite their fearsome appearance these arboreal reptiles still fall prey to certain snakes, and thus need to sleep on thin branches such as this during the night. Thua Thien Hue, Vietnam.
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