Chien C. Lee

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  • Vogel’s Pit Viper (Trimeresurus vogeli). Bach Ma National Park, Vietnam.
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  • Sabah Bamboo Pitviper (Trimeresurus sabahi). Sarawak, Malaysia (Borneo).
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  • Flat-nosed Pitviper (Craspedocephalus puniceus). West Java, Indonesia.
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  • Flat-nosed Pitviper (Craspedocephalus puniceus). West Java, Indonesia.
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  • Black-lipped Shrub Lizard (Pelturagonia nigrilabris). Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Horsfield's Gliding Gecko (Gekko horsfieldii). Sabah, Malaysia (Borneo).
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  • The Bornean Leaf-nosed Pit Viper (Craspedocephalus borneensis) is a well-camouflaged snake, frequently occuring on the forest floor or in low vegetation. They strike readily when disturbed. Sarawak, Malaysia (Borneo).
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  • Triangle Keelback (Xenochrophis trianguligerus). Maliau Basin Conservation Area, Sabah, Malaysia (Borneo).
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  • Kuhl's Gliding Gecko (Gekko kuhli). By means of its large webbed feet and lateral skin flaps, this arboreal gecko is capable of gliding or parachuting between trees. Sarawak, Malaysia (Borneo).
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  • Kuhl's Gliding Gecko (Gekko kuhli). Equipped with webbed feet and parasail-like flaps of skin all along its body, this tree-dwelling gecko is able to ‘parachute’ when leaping through the air to escape predators. This not only softens its landing but also enables it to glide a considerable distance. Sarawak, Malaysia (Borneo).
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  • The rare Kinabalu Brown Pit Viper (Garthius chaseni) is known only from a few mountains in northern Borneo where it occurs in high elevation forest. Very little is known of its life history as it has seldom been observed in the wild. Sabah, Malaysia (Borneo).
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  • The Spiny-headed Shrub Lizard (Pelturagonia spiniceps) is known only from montane forest at a few localities in northwestern Borneo. Pulong Tau National Park, Sarawak, Malaysia (Borneo).
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  • Still wet from a passing rainstorm, a Sabah Bamboo Pitviper (Trimeresurus sabahi) lies in ambush on the side of a tree, waiting for the approach of suitable prey. By maintaining a low metabolism and sometimes remaining in a single position for weeks at a time, tropical vipers such as this can survive by feeding as infrequently as once every few months. Mount Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • Black-lipped Shrub Lizard (Pelturagonia nigrilabris). Sabah, Malaysia (Borneo).
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  • The superbly camouflaged Kinabalu Gliding Gecko (Gekko rhacophorus) is endemic to the montane forests of Mount Kinabalu in northern Borneo.  This poorly-known species is a nocturnal insectivorous tree-dweller and, like other geckos of this genus, possesses skin flaps to assist in a parachuting flight. Sabah, Malaysia (Borneo).
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  • When threatened, the Knob-tailed Gecko (Nephrurus levis) raises its body off the ground to appear larger in size. Cape Range National Park, Western Australia.
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  • White-spotted Cat Snake (Boiga drapiezii), green morph. Sarawak, Malaysia (Borneo).
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  • Few creatures possess a mastery of camouflage the way Madagascar’s leaf-tailed geckos do. This Mossy Leaf-tailed Gecko (Uroplatus sikorae) sleeps by day on a rainforest liana, its outline obscured by frilled dermal flaps that line its body. By always resting in a downward-facing position they may be able to quickly catch unsuspecting insects that climb up from the ground and also conceal their reflective lidless eyes from predators lurking above. Ranomafana National Park, Madagascar.
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  • Madagascar Giant Day Gecko (Phelsuma grandis). Canal des Pangalanes, Madagascar.
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  • Cryptic Chameleon (Calumma crypticum), male. Ranomafana National Park, Madagascar.
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  • The Twin-barred Flying Snake (Chrysopelea pelias) is the smallest and rarest of Southeast Asia's flying snake species. Like other Chrysopelea, this snake hunts lizards and is almost completely arboreal. Sarawak, Malaysia (Borneo).
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  • Although the Banded Coral Snake (Calliophis intestinalis) is not aggressive, it possesses an extremely toxic venom like other Elapids. When disturbed it displays its red tail and the brightly colored bands on its undersurface which serve as a warning signal to would-be predators. Sarawak, Malaysia (Borneo).
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  • Blue-eyed Angle-headed Lizard (Gonocephalus liogaster), male. Sarawak, Malaysia (Borneo).
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  • A juvenile Parson's Chameleon (Calumma parsoni). More frequently occurring in shades of green, this is one of the world's largest chameleons, with adult specimens sometimes reaching the size of a house cat. They are endemic to rainforest habitats on Madagascar and are threatened by both habitat loss and collection for the international pet trade. Andasibe, Madagascar.
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  • Bronzeback (Dendrelaphis terrificus). Gorontalo, Sulawesi, Indonesia.
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  • Spiny-tailed Gecko (Diplodactylus ciliaris). Exmouth, Western Australia.
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  • Hikida's Forest Dragon (Hypsilurus hikidanus). Central Papua, Indonesia (New Guinea).
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  • Sharp-nosed Chameleon (Kinyongia oxyrhina), male. Udzungwa Mountains, Tanzania.
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  • Cloudy Snail-eating Snake (Sibon nebulatus). Limón, Costa Rica.
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  • Bornean Short Python (Python breitensteini). Sarawak, Malaysia (Borneo).
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  • The mimicry of a dead leaf is so precise for the Satanic Leaf-tailed Gecko (Uroplatus phantasticus), they can often hide effectively in plain sight. It has been untested to what degree these geckos actively select a substrate that matches their individual patterns, but U. phantasticus appears to typically hide among dead foliage which suits it perfectly. Ranomafana National Park, Madagascar.
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  • Purple Anole (Anolis purpurescens). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • Malayan Banded Wolf Snake (Lycodon subcinctus), juvenile. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Long-tailed Grass Lizard (Takydromus sexlineatus). Sarawak, Malaysia (Borneo).
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  • Scarcely longer than an earthworm, this juvenile slug-eating snake (Pareas hamptoni) sleeps balanced on top of a fern frond. Many slug-eating snakes retain the  unusual asymmetric dentition of their snail-eating ancestors, an adaptation for pulling snails out of their shells via a process known as ‘mandibular walking’. Bach Ma National Park, Vietnam.
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  • Brown-tailed Racer (Gonyosoma oxycephalum). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Like other small animals of the forest understory, this Borneo Forest Dragon (Gonocephalus bornensis) must juggle his attention between searching for prey (insects and spiders) while keeping a watchful eye out for larger predators. Although this species is unable to glide like the closely-related Dracos, it is nevertheless an agile climber and can move swiftly up and down trees. The impressive mane on the back of this male signifies his maturity - we will readily defend his territory against rival males. Danum Valley, Sabah, Malaysia (Borneo).
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  • Horned Flying Lizard (Draco cornutus), male using his dewlap to display for courtship and territorial aggression. Sabah, Malaysia (Borneo).
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  • Red-headed Reed Snake (Calamaria schlegeli schlegeli), a small soil-dwelling species which feeds on frogs and slugs. Sarawak, Malaysia (Borneo).
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  • Blunthead Tree Snake (Imantodes cenchoa), a lizard hunting specialist that due to its extremely slender body is able to seek out sleeping lizards on even the most delicate of branches. Yasuní National Park, Ecuador.
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  • Although masterful visual predators by day, chameleons lack light-sensitive rods in their eyes and thus have poor night vision. Because of this they typically retire at dusk to a sleeping perch, often returning to a favorite exposed branch which ideally puts them out of the reach of nocturnal snakes. This dozing Two-banded Chameleon (Furcifer balteatus) can be identified as a mature male by the two large rostral horns on his head, ornaments which are used to compete for mates. Endemic to only a small region of Madagascar’s southeastern rainforests, this species has become increasingly endangered and rare due to habitat loss and collection for the pet trade. Ranomafana National Park, Madagascar.
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  • Kirindy Leaf-toed Gecko (Paroedura rennerae), juvenile. Tsingy de Bemaraha National Park, Madagascar.
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  • The Earless Monitor Lizard (Lanthanotus borneensis) is endemic to Borneo where it lives in rocky stream habitats in lowland rainforest. Because of its extreme rarity (it is known from only a few specimens), its life history remains unstudied. It is the only species within its entire family (Lanthanotidae) and was once though to be the missing link between lizards and snakes. Sarawak, Malaysia (Borneo).
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  • Paradise Tree Snake (Chrysopelea paradisi). Sarawak, Malaysia (Borneo).
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  • Wolf snake (Stegonotus sp.). Highland Papua, Indonesia (New Guinea).
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  • Paradise Tree Snake (Chrysopelea paradisi). Sarawak, Malaysia (Borneo).
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  • Lined Pipe Snake (Cylindrophis lineatus) also known as Blanford's Pipe Snake, a rare species endemic to Borneo and known from only a few specimens. When threatened, this species defends itself by raising its tail which is thickened and colored to appear as a head. Sarawak, Malaysia (Borneo).
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  • Carpet Python (Morelia spilota). Queensland, Australia.
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  • A study on perfection in mimicry: a Satanic Leaf-tailed Gecko (Uroplatus phantasticus). One of 14 species of leaf-tailed geckos endemic to Madagascar, this nocturnal insectivore utilizes its incredible camouflage to hide among dead foliage by day. All Uroplatus geckos are under severe threat from deforestation and over-collecting for the international pet trade. Andasibe, Madagascar.
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  • The Bornean Keeled Green Pit Viper (Tropidolaemus subannulatus) is an arboreal species, often laying in wait for prey on the same branch for weeks at a time. Sarawak, Malaysia (Borneo).
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  • Common Sun Skink (Eutropis multifasciata). Sarawak, Malaysia (Borneo).
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  • Four-clawed Gecko (Gehyra mutilata) attending a lanternfly (Pyrops whiteheadi) for secreted honeydew. Host tree is Pometia pinnata. Sabah, Malaysia (Borneo).
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  • When threatened, the harmless Ceylonese Pipe Snake (Cylindrophis maculatus) employs a clever trick: it flattens its tail and raises it to expose the bright checkered pattern underneath. Not only does this divert attacks away from its vulnerable head (which remains hidden beneath), but these bold color patterns resemble venomous coral snakes (Calliophis sp.). Central Province, Sri Lanka.
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  • Common Big-eyed Snake (Mimophis mahfalensis). Toliara, Madagascar.
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  • A juvenile Bornean Dark-necked Slug Snake (Asthenodipsas borneensis). This nocturnal species feeds on slugs and snails in low vegetation. Recently split from A. malaccanus of the Malay Peninsula and Sumatra. Kubah National Park, Sarawak, Malaysia (Borneo).
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  • Ornate Earless Agama (Aphaniotis ornata). Sabah, Malaysia (Borneo).
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  • A species of bent-toed gecko (Cyrtodactylus cf. mikianus) from the Mamberamo Basin in western New Guinea. Papua, Indonesia.
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  • A juvenile Oriental Vine Snake (Ahaetulla prasina) basks in a beam of sunlight in the rainforest of Tangkoko National Park. These highly arboreal snakes are excellent climbers and are adept at hunting lizards, their favored prey. North Sulawesi, Indonesia.
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  • A juvenile Rock Monitor (Varanus albigularis). Mpumalanga, South Africa.
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  • Usambara Eyelash Viper (Atheris ceratophora). Udzungwa Mountains, Tanzania.
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  • Dumeril's Monitor (Varanus dumerilii), juvenile. Sarawak, Malaysia (Borneo).
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  • Banded Malaysian Coralsnake (Calliophis intestinalis). Sarawak, Malaysia (Borneo).
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  • Spiny Ground Gecko (Paroedura bastardi). Amboasary, Madagascar.
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  • This female Madagascar Pygmy Kingfisher (Corythornis madagascariensis) has just been gifted a small lizard (Madascincus melanopleura) by her mate as part of a courtship ritual. Despite their name, these birds are not dependent on water, and typically hunt for insects and small vertebrates on the forest floor. Ranomafana National Park, Madagascar.
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  • Black-headed Cat Snake (Boiga nigriceps). Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Three-banded Bridled Snake (Lycodon tristrigatus). Sarawak, Malaysia (Borneo).
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  • Red-tailed Pipe Snake (Cylindrophis ruffus). Sarawak, Malaysia (Borneo).
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  • Amazon Tree Boa (Corallus hortulanus). Yasuní National Park, Ecuador.
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  • Bornean Keeled Pit Viper (Tropidolaemus subannulatus). Mulu National Park, Sarawak, Malaysia (Borneo).
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  • The Three-striped Ground Snake (Pseudoxyrhopus tritaeniatus) is certainly one of the most beautiful of Madagascan endemic snakes, but why is it colored this way? Bold red stripes are frequently used as an aposematic warning, indicating to other animals of severe danger. It is curious then that not only is this species completely non-venomous but there are in fact no other dangerous snakes in Madagascar that this could possibly be a mimic of. An interesting theory points to the co-occurrence of large red toxic millipedes in the same forests – could these actually be a mimicry model for this snake? On an island legendary for its biological curiosities, nothing seems beyond the realm of possibility. Ranomafana National Park, Madagascar.
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  • Sumatran Pit Viper (Trimeresurus sumatranus). Sarawak, Malaysia (Borneo).
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  • Mossy Leaf-tailed Gecko (Uroplatus sikorae). Mantadia National Park, Madagascar.
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  • Vieira's Snail-eating Snake (Sibon vieirai). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • A sleeping Short-horned Chameleon (Calumma brevicorne) wakes too slowly to react in time when an unaware spider clambers over its face. Andasibe, Madagascar.
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  • The bold coloration of the Red-headed Krait (Bungarus flaviceps) is a warning to would-be predators of the highly potent venom it possesses. This large elapid feeds primarily on other reptiles including lizards and snakes. Sarawak, Malaysia (Borneo).
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  • While pythons undoubtedly hold the crown for the largest and most famous constricting snakes of Indonesia, a single genus of boas also occurs in the rainforests of the eastern archipelago. Often overlooked, the Pacific Keel-scaled Boa (Candoia carinata) seldom exceeds a meter in length. It is an ambush predator of frogs and lizards, and like other boas, bears live young rather than laying eggs. Digul River, South Papua, Indonesia (New Guinea).
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  • Borneo Anglehead Lizard (Gonocephalus bornensis). Sarawak, Malaysia (Borneo).
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  • A Harlequin Gliding Frog (Rhacophorus pardalis) struggles futilely as it is being slowly swallowed alive by a Black-Headed Cat Snake (Boiga nigriceps). Stagnant pools of water in the Borneo rainforest serve as important breeding grounds for many tree frogs which gather around in great numbers, a fact that some predators wisely take advantage of. Kubah National Park, Sarawak, Malaysia (Borneo).
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  • The Bornean Forest Dragon (Gonocephalus bornensis) is a large arboreal agamid lizard endemic to the rainforests of Borneo. It feeds on ants and spiders. Sarawak, Malaysia (Borneo).
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  • The Oriental Vine Snake (Ahaetulla prasina) is an agile climber, feeding on small birds and lizards with its excellent binocular vision. Sabah, Malaysia (Borneo).
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  • Two-banded Chameleon (Furcifer balteatus), female. Ranomafana National Park, Madagascar.
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  • Bornean Keeled Green Pit Viper (Tropidolaemus subannulatus). Sarawak, Malaysia (Borneo).
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  • A Pilbara Death Adder (Acanthophis wellsi) from the desert regions of Western Australia. Although superficially resembling vipers by their stout body and habit of ambushing prey, death adders are actually elapids and more closely related to cobras, kraits, and coral snakes. Cape Range National Park, Western Australia.
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  • A young Parson's Chameleon (Calumma parsonii) in the rainforest of Masoala National Park. Antsiranana, Madagascar.
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  • Green Tree Python (Morelia azurea), juvenile.
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  • Canopy Slug-eating Snake (Sibon canopy), a new species described in January 2023, endemic to Panama. Panamá Oeste, Panama.
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  • With colors that seem to glow with an unearthly radiance, a male Osa Anole (Anolis osa) betrays his normally cryptic appearance to briefly flash his extended dewlap. Anole lizards are famous for these flamboyant visual signals – they are used to attract mates, ward off rivals, and even sometimes dissuade potential predators. A 2015 study found that the translucent quality of anole dewlaps was a crucial component in their ability to transmit these color signals effectively, particularly in shady environments; under the right conditions they possess an almost unnatural luminance. This species, which is endemic to Costa Rica’s Osa Peninsula, spends most of its time on perches near ground level in the dappled light of the rainforest understory. Osa Peninsula, Puntarenas, Costa Rica.
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  • With night vision hundreds of times more sensitive than ours and the ability to discern color even in near total darkness, Madagascar’s leaf-tailed geckos (Uroplatus spp.) are superbly adapted as nocturnal insect hunters. Large eyes, however, demand special maintenance. Like most geckos, Uroplatus have no eyelids and cannot blink; instead, they utilize their long tongues like windshield wipers, keeping their eyes moist and clean. This is a young male Satanic Leaf-tailed Gecko (U. phantasticus), a denizen of the island’s rainforest zone and a superb mimic of dead foliage. Ranomafana National Park, Madagascar.
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  • The Black-headed Cat Snake (Boiga nigriceps) is an arboreal predator of birds and other snakes. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Neotropical Snail-Eater (Dipsas indica). Yasuní National Park, Ecuador.
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  • The mimicry of a dead leaf is so precise for the Satanic Leaf-tailed Gecko (Uroplatus phantasticus), they can often hide effectively in plain sight. It has been untested to what degree these geckos actively select a substrate that matches their individual patterns, but U. phantasticus appears to typically hide among dead foliage which suits it perfectly. Ranomafana National Park, Madagascar.
    cld1916615.jpg
  • Smith's Green-eyed Gecko (Gekko smithii). Kinabatangan River, Sabah, Malaysia (Borneo).
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  • Inger's Slug-eating Snake (Asthenodipsas ingeri). Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • Males of the Madagascar's Satanic Leaf-tailed Gecko (Uroplatus phantasticus) can usually be distinguished by their elaborately notched tails, a trait which gives them better mimicry of a leaf that has been partially eaten by insects. Exactly why this is a sexually dimorphic feature in these geckos is unknown, but in some animals different camouflage patterns in males and females can arise where each sex spends their time hiding in different locations and thus face differing selective pressures on their appearance. Ranomafana National Park, Madagascar.
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  • Like most snail-eating snakes, the Blunt-headed Tree Snake (Aplopeltura boa) is small and slender with an abnormally large head. They possess an asymmetric arrangement of teeth in their lower jaw which enables them to pry snails from their shells (most of which swirl in a clockwise direction) by a process referred to as "mandible walking". This species is widespread in Southeast Asia and ranges from Myanmar to the Philippines and Sunda islands. Sabah, Malaysia (Borneo).
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  • Spotted Cat Snake (Boiga drapiezii). Sarawak, Malaysia (Borneo).
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  • Bornean Angle-headed Lizard (Gonocephalus bornensis), male. Sarawak, Malaysia (Borneo).
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  • Sabah Keelback (Hebius flavifrons). Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Komodo Dragons (Varanus komodoensis), like all monitor lizards, posses forked tongues which are used to smell the air (chemoreception) in a similar fashion to snakes. Rinca Island, East Nusa Tenggara, Indonesia.
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  • One of the most dangerous snakes in New Guinea: the Smooth-scaled Death Adder (Acanthophis laevis). Like vipers, these snakes are thick-bodied with a wide triangular head and are relatively slow moving, preferring to lie in wait for suitable prey to venture within striking distance. However, their resemblance to vipers (and true adders) is only superficial - they are actually members of the family Elapidae, which includes cobras, kraits, and coral snakes. Papua, Indonesia (New Guinea).
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