Chien C. Lee

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  • Asian Toad (Duttaphrynus melanostictus), an introduced and invasive species in New Guinea. Nimbokrang, Papua, Indonesia (New Guinea).
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  • The Asian Eyebrow-ridge Toad (Duttaphrynus melanostictus) is a common species widespread in South and Southeast Asia.  It has a distinctive large parotoid glands behind the head, which are known to secrete a toxic substance when the toad is disturbed.
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  • Borneo Horned Frog (Pelobatrachus nasutus), female, with Matang Narrow-mouthed Frog (Microhyla nepenthicola) - comparison between one of the largest and smallest frogs in Borneo. Sarawak, Malaysia (Borneo).
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  • Endemic to the mountains of northern Borneo, the Kinabalu Horned Frog (Pelobatrachus baluensis) breeds only in clear rocky streams with cold water. By day they lie camouflaged among the leaf litter on the forest floor, but they emerge at night to call for mates. Sabah, Malaysia (Borneo).
    cld1802598.jpg
  • Bornean Horned Frog (Pelobatrachus nasutus). Found throughout the lowland rainforests of Borneo, the loud honking call of this frog is commonly heard before the onset of heavy thunderstorms. When disturbed it remains completely motionless, relying on its excellent camouflage to escape detection from potential predators. Sarawak, Malaysia (Borneo).
    cld1704491.jpg
  • Smith's Litter Frog (Leptobrachium smithi). Krabi, Thailand.
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  • Borneo Horned Frog (Pelobatrachus nasutus). Sabah, Malaysia (Borneo).
    cld1409666.jpg
  • With half a dozen species, Borneo is famous for its horned frogs. These are denizens of the rainforest floor and as such are expertly camouflaged for hiding among leaf litter. Most species rely so much on crypsis to avoid detection they have neglected other adaptations to escape predators, such as having weak legs for jumping. This is the rarest of the Bornean species: the Mulu Horned Frog (Megophrys dringi) which has only been known from a handful of sightings. This paucity of records is undoubtedly due to the remote locality where it occurs: high-elevation moss forests adjacent to clear mountain streams. Mulu National Park, Sarawak, Malaysia (Borneo).
    cld2006095.jpg
  • Bornean Horned Frog (Pelobatrachus nasutus). Found throughout the lowland rainforests of Borneo, the loud honking call of this frog is commonly heard before the onset of heavy thunderstorms. When disturbed it remains completely motionless, relying on its excellent camouflage to escape detection from potential predators. Sarawak, Malaysia (Borneo).
    cld1704562.jpg
  • The rare Montane Horned Frog (Pelobatrachus kobayashii) is endemic to mountain rainforests of Sabah in northern Borneo. Like other members of the genus, this large terrestrial frog relies on its superb camouflage to evade detection from predators. Sabah, Malaysia (Borneo).
    cld1303455.jpg
  • Endemic to the mountains of northern Borneo, the Kinabalu Horned Frog (Pelobatrachus baluensis) breeds only in clear rocky streams with cold water. By day they lie camouflaged among the leaf litter on the forest floor, but they emerge at night to call for mates. Sabah, Malaysia (Borneo).
    cld1605084.jpg
  • With half a dozen species, Borneo is famous for its horned frogs. These are denizens of the rainforest floor and as such are expertly camouflaged for hiding among leaf litter. Most species rely so much on crypsis to avoid detection they have neglected other adaptations to escape predators, such as having weak legs for jumping. This is the rarest of the Bornean species: the Mulu Horned Frog (Megophrys dringi) which has only been known from a handful of sightings. This paucity of records is undoubtedly due to the remote locality where it occurs: high-elevation moss forests adjacent to clear mountain streams. Mulu National Park, Sarawak, Malaysia (Borneo).
    cld2006086.jpg
  • Inger's Black-eyed  Litter Frog (Leptobrachium ingeri). Sarawak, Malaysia (Borneo).
    cld1823839.jpg
  • The rare Montane Horned Frog (Pelobatrachus kobayashii) is endemic to mountain rainforests of Sabah in northern Borneo. Like other members of the genus, this large terrestrial frog relies on its superb camouflage to evade detection from predators. Sabah, Malaysia (Borneo).
    cld1802569.jpg
  • Dring's Slender Litter Frog (Leptolalax dringi). Sarawak, Malaysia (Borneo).
    cld1602459.jpg
  • The Mindanao Horned Frog (Pelobatrachus stejnegeri) is endemic to the rainforests of the south-eastern Philippines where it is threatened by habitat loss. Mindanao, Philippines.
    cld1202723.jpg
  • Found throughout the lowland rainforests of Sarawak, the loud honking call of the Bornean Horned Frog (Pelobatrachus nasutus) is commonly heard before the onset of heavy thunderstorms. When disturbed it remains completely motionless, relying on its excellent camouflage to escape detection from potential predators. Sabah, Malaysia (Borneo).
    cld08121117.jpg
  • Bornean Horned Frog (Pelobatrachus nasutus), vocalising. Sarawak, Malaysia (Borneo).
    cld1500932.jpg
  • Found throughout the lowland rainforests of Sarawak, the loud honking call of the Bornean Horned Frog (Pelobatrachus nasutus) is commonly heard before the onset of heavy thunderstorms.  When disturbed it remains completely motionless, relying on its excellent camouflage to escape detection from potential predators. Mulu National Park, Sarawak, Malaysia (Borneo).
    cld06070437.jpg
  • The rare and little known Rough Horned Frog (Pelobatrachus edwardinae) is endemic to Borneo. Sarawak, Malaysia (Borneo).
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  • Rough Horned Frog (Pelobatrachus edwardinae), male. Sarawak, Malaysia (Borneo).
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  • Found throughout the lowland rainforests of Sarawak, the loud honking call of the Bornean Horned Frog (Pelobatrachus nasutus) is commonly heard before the onset of heavy thunderstorms.  When disturbed it remains completely motionless, relying on its excellent camouflage to escape detection from potential predators. Mulu National Park, Sarawak, Malaysia (Borneo).
    cld06070439.jpg
  • Brown Bullfrog (Kaloula baleata), male. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Brown Bullfrog (Kaloula baleata). Sabah, Malaysia (Borneo).
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  • Banded Bullfrog (Kaloula pulchra), an introduced frog to Borneo. Sarawak, Malaysia.
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