Chien C. Lee

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  • Like some kind of karmic balance, it has often been remarked that the more plain-looking a bird is, the more beautiful its song, whereas those bestowed with gorgeous plumage tend to have simple calls. Research across hundreds of bird species has revealed a negative correlation between these traits, suggesting that there may be a trade-off in visual vs. acoustic signals when it comes to attracting mates. Likely falling at one end of this spectrum, Birds-of-paradise certainly are among the most extravagantly ornamented species in the world, but their highly varied calls often consist of harsh squawks and penetrating whistles, which although extremely loud are anything but melodious. Here, a male Wilson’s BoP (Cicinnurus respublica) raucously advertises his presence above his display ground, though if a female approaches he will fall silent and rely on his feathers for the real game of courtship. Waigeo Island, West Papua, Indonesia.
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  • The Magnificent Bird of Paradise (Cicinnurus magnificus) is restricted to New Guinea, where it occurs throughout the island in suitable hill forest. Male birds clear a personal display site near the forest floor where they put on an impressive display of their vibrant plumage for visiting females. West Papua, Indonesia (New Guinea).
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  • The Magnificent Bird of Paradise (Cicinnurus magnificus) is restricted to New Guinea, where it occurs throughout the island in suitable hill forest. Male birds clear a personal display site near the forest floor where they put on an impressive display of their vibrant plumage for visiting females. West Papua, Indonesia (New Guinea).
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  • With his otherworldly plumage a male Wilson's Bird-of-paradise (Cicinnurus respublica) shows off his gaudy colors at a dancing ground he has cleared on the forest floor. Endemic to only two islands off the western coast of New Guinea, Wilson's BoP is surely one of the most bizarre in a family of birds already renowned for their exotic appearances.
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  • Wilson's Bird-of-paradise (Diphyllodes respublica), male courting female at his display ground. West Papua, Indonesia (New Guinea)
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  • The incredible vibrance of many Birds-of-paradise is in part due to the deep black feathers which make their more colorful plumage simply ‘pop’. A recent study looking closely at these black feathers found that they had a unique nanostructure that is super-efficient in light absorption. This results in one of the darkest blacks found in nature, rivaling those of highly specialized synthetic materials. Here, a Vogelkop Superb Bird-of-paradise (Lophorina niedda) waits at his display (a fallen log) for the arrival of a female. West Papua, Indonesia (New Guinea).
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  • Magnificent Bird-of-paradise (Diphyllodes magnificus), male calling from his display ground, showing the bright yellow inside of his mouth. West Papua, Indonesia (New Guinea).
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  • Transforming into an extravagant bouquet of colors and shapes, a male Magnificent Bird-of-paradise (Diphyllodes magnificus) puts on his best show in the hopes of impressing a female visitor to his courting ground. She has a critical eye for imperfections as she will be selecting him based on the merits of his display alone – a testament to his overall fitness and desirable genes. This selective pressure is the driving force behind the evolution of some of the most incredible avian spectacles in the world.
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  • It’s impossible for a single image to capture all of the multifaceted ornamentation of a male Magnificent Bird-of-paradise (Cicinnurus magnificus). Depending on the viewpoint and which of his assets he chooses to flaunt, his display can range from a brilliant metallic green breast plate to a golden yellow nape fan. The remarkable curled tail wires, which often appear black in color, flash an incredible turquoise when the angle is just right – a tool he often uses to bedazzle female visitors to his courting ground. Arfak Mountains, West Papua, Indonesia (New Guinea).
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  • A Red-bearded Bee-eater (Nyctyornis amictus) living up to its name: here with a large black carpenter bee (Xylocopa sp.) it has caught in mid-air. While most bee-eaters prefer open woodlands and forest edges, this species is unusual in that it is typically found deep within rainforest. Flashy colors in birds are often possessed only by the males and typically function as a means of advertising for a mate. However, in this species both males and females sport a bright red face, which suggests that the coloration may serve a different purpose. One theory is that this may serve to attract flower-feeding insects to venture closer for a look - a convenient way for the bird to catch an easy meal. Danum Valley, Sabah, Malaysia (Borneo).
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  • The rare and elusive Gurney's Pitta (Pitta gurneyi) is regarded as one of the world's most endangered birds.  Until recently this species was known from less than 20 individuals all occurring in a single reserve in southern Thailand.  Following the discovery of additional populations in southern Myanmar in 2003, the status Gurney's Pitta was downgraded from Critically Endangered to Endangered.  This species occurs only in flat lowland rainforest, a rapidly disappearing habitat in this region due to logging and oil-palm plantations. The adult male bird pictured here sports a leg band from a recent population survey. Krabi, Thailand.
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  • Endemic to northern Borneo, the Black-and-crimson Pitta (Pitta ussheri) is a shy forest bird which feeds on worms and other terrestrial invertebrates. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • The Palawan Peacock Pheasant (Polyplectron napoleonis) is a shy ground-dwelling bird endemic to the rainforests the island of Palawan, Philippines.
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  • The shy Maleo (Macrocephalon maleo) is a large megapode bird endemic to Sulawesi. They do not build nests but instead bury their eggs in huge pits dug in hot volcanic sand. The precocious young are able to fly and fend for themselves immediately after hatching. Central Sulawesi, Indonesia.
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  • The Rufous Piculet (Sasia abnormis) is the smallest woodpecker in Borneo.  Here a bird is making its territorial call by drumming on a spiny rattan petiole. Sabah, Malaysia (Borneo).
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  • The Black-naped Monarch (Hypothymis azurea) is an active insect-eating bird of the forest canopy. Males, such as the one pictured here, sport a richer blue plumage than the greyish-colored females. Sarawak, Malaysia (Borneo).
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  • A nesting Common Potoo (Nyctibius griseus) in the dense mangrove forest of Colombia’s Utría National Park. Rather than construct a nest, the bird deposits its single egg in a small depression along a branch. Active only by night, potoos sit motionless during the day, using their incredible camouflage to remain hidden. Choco, Colombia.
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  • The Diabolical Nightjar (Eurostopodus diabolicus) produces such an eerie call that local people in Sulawesi hold the superstitious belief that it is a demonic entity, hence its name. The life history of this rare bird is very poorly known as until recently only a single specimen had ever been found. Like other nightjars it rests by day on the ground, using its superb camouflage to escape detection from predators. Central Sulawesi, Indonesia.
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  • At only 12cm in length, the Sulawesi Dwarf Kingfisher (Ceyx fallax) is one of the smallest kingfishers. It is a bird of the forest, feeding on small lizards and insects, and is endemic to Sulawesi and a few offshore islands, although scarce and patchily distributed. Gorontalo, Indonesia.
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  • Although the vast Australian outback is famous for its aridity, poor soils, and tortuous heat in summer, a surprising number of colorful creatures make this habitat their home. One of these is the aptly named Splendid Fairy-wren (Malurus splendens), a dainty yet bold bird that lives in small groups that hunt insects among the desert scrub. Despite the male’s gaudy plumage (as shown here), he will supplement his courtship efforts by wooing the female with pink or purple flower petals he has carefully plucked. Shark's Bay, Western Australia.
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  • The Great Eared Nightjar (Lyncornis macrotis) is one of the largest in its family, reaching lengths of over 40cm. Like other nightjars, this cryptically-colored nocturnal bird relies on its camouflage to remain undetected while sleeping on or near the forest floor during the day. North Sulawesi, Indonesia.
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  • At only 12cm in length, the Sulawesi Dwarf Kingfisher (Ceyx fallax) is one of the smallest kingfishers. It is a bird of the forest, feeding on small lizards and insects, and is endemic to Sulawesi and a few offshore islands, although scarce and patchily distributed. Gorontalo, Sulawesi, Indonesia.
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  • The Palawan Peacock Pheasant (Polyplectron napoleonis) is a shy ground-dwelling bird endemic to the rainforests the island of Palawan, Philippines.
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  • Masked Bowerbird (Sericulus aureus) male repairing his bower after it was destroyed by a rival bird. West Papua, Indonesia (New Guinea).
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  • Pygmy Longbill (Oedistoma pygmaeum), a nectar-feeding bird showing convergent evolution with sunbirds and honeyeaters, but part of the New Guinean endemic family Melanocharitidae. West Papua, Indonesia (New Guinea).
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  • With a flair for home decor, the Vogelkop Bowerbird (Amblyornis inornata) produces perhaps the most artful creation of any animal, all in the name of romance. Here, a male bird shows off his collection of freshly collected flowers, fungi, beetle shells, and blue berries, neatly organized in piles laid out on a mat of meticulously cleaned moss. The twig hut in the back is not a nest - rather its construction is made entirely for the purpose of impressing a visiting female who will choose to mate with him only after a thorough inspection of his display. Arfak Mountains, West Papua, Indonesia (New Guinea).
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  • A shy bird of the rainforest floor, the beautiful Blue-headed Pitta (Hydrornis baudii) is endemic to Borneo. Danum Valley Conservation Area, Sabah, Malaysia.
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  • The Oriental Bay-Owl (Phodilus badius) is a denizen of dense rainforest; with its short and rounded wings it is able to manoeuvre between branches and twisted understory lianas while hunting for small animals, often stopping to perch peculiarly on vertical saplings as seen here. Like its larger relatives, the barn owls, the concave shape of the face is designed to focus sounds towards its ears, enabling the bird to hone in on even the faintest sounds of prey. Sarawak, Malaysia (Borneo).
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  • Red-crowned Barbet (Psilopogon rafflesii), a frugivorous bird from the rainforests of Southeast Asia. Sarawak, Malaysia (Borneo).
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  • The handsomely-proportioned Short-legged Ground-roller (Brachypteracias leptosomus) belongs to yet another of Madagascar’s endemic bird families, Brachypteraciidae. Patchily distributed in the rainforests of eastern Madagascar, it is a seldom-seen species owing to its habit of perching completely still and quiet for long periods of time in the lower canopy. They feed on large insects and other invertebrates. Ranomafana National Park, Madagascar.
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  • Known from only a handful of sightings and photos, the elusive Spectacled Flowerpecker (Dicaeum dayakorum) is the first new endemic bird discovered in Borneo in over 100 years.
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  • Once considered one of the three most elusive birds in the world, the Long-tailed Ground Roller (Uratelornis chimaera) is a prized sighting for any birdwatcher. The entire family of Ground Rollers (Brachypteraciidae, 5 species) is endemic to Madagascr, but while most are inhabitants of rainforest, this is the only species found in in the arid spiny desert of the island’s extreme southwest. Even within this region it has as very restricted distribution and occurs at low densities, with only a few birds per square kilometer. Adapted for running swiftly on the ground with their powerful legs, these birds hunt for insects and small animals in the sand and leaf-litter. Toliara, Madagascar.
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  • Only the male Wallace's Standardwing (Semioptera wallacii) possesses the highly ornamental plumage, females being plain brown in coloration. This species is endemic to Halmahera and a few offshore islands and is one of the westernmost birds of paradise (family Paradisaeidae). Halmahera, North Maluku, Indonesia.
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  • Magnificent Riflebird (Lophorina magnifica), male calling from display perch. Heard far more often than seen, these uncommon and extremely shy birds are responsible for the most quintessential sound of the Papuan rainforest. Papua, Indonesia (New Guinea).
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  • The bizarre and rare Bornean Bristlehead (Pityriasis gymnocephala) is one of the most enigmatic birds of Borneo and is highly sought after by birdwatchers. It is the only member of its family and its nearest relations to other birds has been highly debated. Bristleheads are found in small nomadic flocks traveling in the middle to upper canopy where they feed on insects. Sabah, Malaysia (Borneo).
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  • Looking like it might have flown straight out of the Jurassic, the rare and reclusive Horned Guan (Oreophasis derbianus) possesses one of the most remarkable head ornaments of all birds. They live only high in the cloud forests of Chiapas and Guatemala and over the past hundred years have suffered greatly from habitat loss and hunting, with some current surveys estimating at fewer than 1000 birds remaining in the wild. El Triunfo Biosphere Reserve, Chiapas, Mexico.
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  • The bizarre and rare Bornean Bristlehead (Pityriasis gymnocephala) is one of the most enigmatic birds of Borneo and is highly sought after by birdwatchers. It is the only member of its family and its nearest relations to other birds has been highly debated. Bristleheads are found in small nomadic flocks traveling in the middle to upper canopy where they feed on insects. Sabah, Malaysia.
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  • Only the male Wallace's Standardwing (Semioptera wallacii) possesses the highly ornamental plumage, females being plain brown in coloration. This species is endemic to Halmahera and a few offshore islands and is one of the westernmost birds of paradise (family Paradisaeidae). Halmahera, Indonesia.
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  • Almost indistinguishable from the branch on which it rests, a Papuan Frogmouth (Podargus papuensis) perches motionless next to its chick. Like other frogmouths, these birds scarcely build any nest at all and rely on camouflage to escape the detection of predators. Papua, Indonesia (New Guinea).
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  • This female Madagascar Pygmy Kingfisher (Corythornis madagascariensis) has just been gifted a small lizard (Madascincus melanopleura) by her mate as part of a courtship ritual. Despite their name, these birds are not dependent on water, and typically hunt for insects and small vertebrates on the forest floor. Ranomafana National Park, Madagascar.
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  • A lone Scarlet Macaw (Ara macao) ventures down to the rainforest floor to take a sip of mineral mud at a clay lick. This is a common phenomenon with the larger parrots of the region and was often believed to be due to the fact that the birds ingest the clay to help neutralize toxins in the fruits that they consume. More recent observations suggest that they may simply be obtaining salt – a mineral that is in particularly scarce supply in the western Amazon Basin. Yasuní National Park, Ecuador.
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  • At the onset of dusk, a pair of Many-banded Aracari (Pteroglossus pluricinctus) arrive at their roosting tree hole where they will be joined by the rest of their family group to spend the night. These birds are unusual among toucans for their gregarious behavior with as many as half a dozen sleeping together. Yasuní National Park, Ecuador.
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  • Ubiquitous throughout the forests of the island, the Madagascar Crested Drongo (Dicrurus forficatus) is well-respected by other birds for its pugnacious and aggressive attitude. They are extremely agile on the wing, swooping out from perches to snatch insects in mid-air, and have even been known to take prey as large as small chameleons and geckos. Ranomafana National Park, Madagascar.
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  • A male Whiskered Treeswift (Hemiprocne comata). Unlike true swifts, these birds can often be seen perching on exposed branches from which they make short sallying flights after insects. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • A jewel of Ecuador’s high-elevation cloud forests is the spectacular Plate-billed Mountain Toucan (Andigena laminirostris). Unlike the more familiar toucans of lowland rainforests, these birds inhabit the coldest and wettest mountains of the Andes. Despite their predilection for these remote habitats, they are threatened by both habitat loss and capture for the pet trade. Mindo, Ecuador.
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  • A male Great Argus Pheasant (Argusianus argus) is one of the most magnificent birds in the Bornean rainforest. Despite its large size it is shy and seldom seen, though its haunting call is frequently heard. Sabah, Malaysia (Borneo).
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  • A Dusky Broadbill (Corydon sumatranus) carrying nesting material. This species travels in semi-nomadic family groups and up to 20 individual birds have been observed contributing to the construction of a single nest. Sabah, Malaysia (Borneo).
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  • A Malagasy Kingfisher (Corythornis vintsioides), found only on Madagascar and the neighboring islands of Comoros. Although famous for its lemurs, well over 100 species of birds are endemic to Madagascar, including several entire families. Tomasina, Madagascar.
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  • A carefully arranged pile of Aeschynanthus flowers forms the centerpiece of a Vogelkop Bowerbird's (Amblyornis inornata) display. The elaborately constructed twig hut with various colorful ornaments (here including flowers, beetle elytra, and fruit husks) is constructed by the male solely for the purpose of impressing female birds to select him as a mate. Each male will use their own artistic preferences in selecting various colors for their display, and theft of highly desirable items by competitors is a frequent occurrence. West Papua, Indonesia (New Guinea).
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  • A Feline Owlet-nightjar (Aegotheles insignis) rests in a dense thicket of vegetation by day. This unusual family of nocturnal insectivorous birds, whose center of distribution is in New Guinea, is neither closely related to owls nor nightjars, and are placed in their own order. West Papua, Indonesia (New Guinea).
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  • Yellow-bellied Sunbird-asity (Neodrepanis hypoxantha), male in breeding plumage. Despite their appearance, these birds are closely related to Asian broadbills; their slender beaks representing an example of convergence with sunbirds due to their nectar-feeding habits. Ranomafana National Park, Madagascar.
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  • Endemic to the mossy cloud forests of Papua, a Mountain Owlet-nightjar (Aegotheles albertisi) rests by the day in a thicket of vegetation. Biologists once believed that these strange cryptic birds (family Aegothelidae) were close relatives of the frogmouths due to their similar appearance and habits. Studies of their skulls has revealed that they are in fact more closely related to hummingbirds and swifts. Almost the entire family (7 of 10 species) is restricted to New Guinea. Arfak Mountains, West Papua, Indonesia.
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  • Short-legged Ground Roller (Brachypteracias leptosomus), one of several families of birds which are completely endemic to Madagascar. Mantadia National Park, Madagascar.
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  • Yellow-bellied Sunbird-asity (Neodrepanis hypoxantha), male in breeding plumage. Despite their appearance, these birds are closely related to Asian broadbills; their slender beaks representing an example of convergence with sunbirds due to their nectar-feeding habits. Ranomafana National Park, Madagascar.
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  • In contrast with its rather unremarkable appearance, the Sapayoa (Sapayoa aenigma) is one of South America's most enigmatic birds because it has no known relatives in the neotropics. Genetic studies have placed it close to the Asian broadbills (Eurylaimidae). Utría National Natural Park, Choco, Colombia.
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  • With his crop full of food and water, a male Plain-pouched Hornbill (Rhyticeros subruficollis) returns to his nest in the cavity of a tree wherein the female waits sealed inside. During the nesting period, which can last several months, his mate will be completely dependent on his regular visits to provide her with everything she needs while she incubates the young and raises the chicks. Unfortunately, his duties on this day have been complicated by the annoying attentions of a Greater Racket-tailed Drongo (Dicrurus paradiseus) which has perhaps also been nesting nearby. Although much smaller than the hornbill, drongos are known for their bold and aggressive behavior, especially towards other birds that could be potential predators of their nests. Huai Kha Khaeng Wildlife Sanctuary, Thailand.
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  • Red-bearded Bee-Eater (Nyctyornis amictus), male. Sabah, Malaysia (Borneo).
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  • Extremely agile on the wing, the Bat Hawk (Macheiramphus alcinus) is a specialist predator of bats and swiftlets, often waiting near cave entrances for its prey to emerge. Sarawak, Malaysia (Borneo).
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  • Crested Serpent Eagle (Spilornis cheela). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Once considered a subspecies of the widespread Crested Serpent Eagle (Spilornis cheela), the Sulawesi Serpent Eagle has a distinctive plumage and is endemic to Sulawesi. Like other serpent eagles it feeds on snakes, lizards, and occasional small mammals. Central Sulawesi, Indonesia.
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  • Red-headed Barbet (Eubucco bourcierii), male. Mindo, Ecuador.
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  • With tail feathers nearly a meter long, a male Black Sicklebill (Epimachus fastosus) struts his incredible plumage on a display perch high in the Arfak Mountains of West Papua. One of the largest of all the birds-of-paradise, this shy sicklebill is found only in remote cloud forests on the island of New Guinea. Although normally keeping to the tree tops, the male carefully selects an exposed perch lower in the canopy from where he will call and display for females at dawn.
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  • A male Whitehead's Broadbill (Calyptomena whiteheadi) returns to its nest where it passes food to the hungry chick. Both male and female care for the nestlings, providing insects and fruit, as well as removing fecal pellets. This scarcely-seen species is the largest of all the green broadbills (reaching 27cm in length) and is endemic to the montane forests of Borneo. Sabah, Malaysia.
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  • This Collared Nightjar (Gactornis enarratus), a Madagascar endemic, hides itself among the leaf litter of the forest floor while it sleeps. Placed in its own genus, this species is distinct from all other nightjars (family Caprimulgidae) by its plumage, eggs, and behavior. Mantadia National Park, Madagascar.
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  • The Black-sided (or Bornean) Flowerpecker (Dicaeum monticolum) is the only montane flowerpecker in Borneo and is endemic to the island. Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • Temminck's Sunbird (Aethopyga temminckii), male. Sabah, Malaysia (Borneo).
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  • Sunda Frogmouth (Batrachostomus cornutus), female. Sabah, Malaysia (Borneo).
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  • Plain-pouched Hornbill (Rhyticeros subruficollis), male at nest. Huai Kha Khaeng Wildlife Sanctuary, Thailand.
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  • Pitta-like Ground-roller (Atelornis pittoides) with earthworm. Ranomafana National Park, Madagascar.
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  • Guatemalan Yellow Grosbeak (Pheucticus chrysopeplus aurantiacus), vocalising. El Triunfo Biosphere Reserve, Chiapas, Mexico.
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  • Long-tailed Sylph (Aglaiocercus kingii). Papallacta, Ecuador.
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  • Madagascar Plover (Charadrius thoracicus). Toliara, Madagascar.
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  • Borneo Necklaced Partridge (Tropicoperdix graydoni), considered by some to be a subspecies of Chestnut-necklaced Partridge (Tropicoperdix charltonii) from mainland Southeast Asia. Sabah, Malaysia (Borneo).
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  • Brehm's Tiger-parrot (Psittacella brehmii). West Papua, Indonesia (New Guinea).
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  • Madagascan Nightjar (Caprimulgus madagascariensis). Tomasina, Madagascar.
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  • Endemic to the mountains of Borneo, the Chestnut-hooded Laughingthrush (Pterorhinus treacheri) often travels in noisy flocks while they forage for insects and fruit. Sabah, Malaysia (Borneo).
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  • Scarcely larger than your thumb, the Buff-faced Pygmy Parrot (Micropsitta pusio) is the world’s smallest parrot. Here, a mated pair peer out from their nest hole that they have excavated in an arboreal termite mound within the lowland rainforest of western New Guinea. Papua, Indonesia.
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  • Although occurring widely in Borneo, Sumatra, and Peninsular Malaysia, the Striped Wren-babbler (Kenopia striata) is nowhere abundant and is considered Near Threatened due to habitat loss. Inhabiting lowland forests, its presence is often first confirmed by its loud clear and persistent whistling calls. Sabah, Malaysia (Borneo).
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  • The seldom seen Archbold's Nightjar (Eurostopodus archboldi) is endemic to the highest mountains of New Guinea, where it can be found in open alpine habitats. Like other nightjars it sleeps by day on the ground using excellent camouflage to conceal itself. West Papua, Indonesia (New Guinea).
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  • The enormous Palm Cockatoo (Probosciger aterrimus), sometimes called the Goliath Cockatoo, is one of the largest parrots in Asia. It is found in the rainforests of New Guinea and northern Australia. West Papua, Indonesia (New Guinea).
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  • Bornean Frogmouth (Batrachostomus mixtus). Sarawak, Malaysia (Borneo).
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  • White-bellied Woodstar (Chaetocercus mulsant). Papallacta, Ecuador.
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  • Swallow-tailed Nightjar (Uropsalis segmentata). Parque Natural Chicaque, Cundinamarca, Colombia.
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  • Ecuadorian Hillstar (Oreotrochilus chimborazo), female feeding juvenile female. Cotopaxi National Park, Ecuador.
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  • Barred Eagle-owl (Ketupa sumatrana). Danum Valley, Sabah, Malaysia (Borneo).
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  • Malagasy Kestrel (Falco newtoni). Akanin'ny Nofy, Madagascar.
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  • Masked Trogon (Trogon personatus), male. Mindo, Ecuador.
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  • Endemic to Sulawesi and the Sula Islands, the Sulawesi Black Pigeon (Turacoena manadensis) inhabits coastal, lowland, and hill forests. Gorontalo, Indonesia.
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  • Green Honeycreeper (Chlorophanes spiza), male. Colón, Panama.
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  • Blue-banded Pitta (Erythropitta arquata). Sarawak, Malaysia (Borneo).
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  • Little Green-Pigeon (Treron olax), male feeding on fruits of Glochidion rubrum. Sabah, Malaysia (Borneo).
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  • Blue-banded Pitta (Erythropitta arquata). Sarawak, Malaysia (Borneo).
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  • Western Crowned-pigeon (Goura cristata). Waigeo Island, West Papua, Indonesia.
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  • Minahasa Hooded Pitta (Pitta sordida forsteni). North Sulawesi, Indonesia.
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  • Olive-backed Sunbird (Nectarinia jugularis frenata), female at nest. Halmahera, Indonesia.
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  • Velvet Asity (Philepitta castanea), male. Ranomafana National Park, Madagascar.
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  • Malagasy Green Sunbird (Cinnyris notatus) feeding at flowers of Ravenala grandis. Ranomafana National Park, Madagascar.
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  • Rhinoceros Hornbill (Buceros rhinoceros). Sepilok Forest Reserve, Sabah, Malaysia (Borneo).
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  • Rufous-headed Ground-roller (Atelornis crossleyi). Ranomafana National Park, Madagascar.
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  • Bushy-crested Hornbill (Anorrhinus galeritus), juvenile. Sepilok Forest Reserve, Sabah, Malaysia (Borneo).
    cld1930460.jpg
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