Chien C. Lee

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  • Earth-boring dung beetle (Blackburnium rhinoceros), male. This unusual dung beetle is a member of the family Geotrupidae, often referred to as the earth-boring dung beetles. They are known for laying their eggs in burrows which are sometimes provisioned with leaf litter or dung. Wasur National Park, South Papua, Indonesia (New Guinea).
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  • Dung beetle (Paragymnopleurus sparsus), rolling scat of Sambar Deer. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Dung beetle (Scarabaeus sp.), rolling dung. Bundala National Park, Sri Lanka.
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  • Dung beetle (Proagoderus watanabei), male feeding on herbivore scat. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • This small dung beetle (Paragymnopleurus maurus) is preparing a ball made from monkey droppings in which it will deposit its eggs and then bury. Sarawak, Malaysia (Borneo).
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  • Dung beetle (Paragymnopleurus maurus), rolling scat of Sambar Deer. Danum Valley, Sabah, Malaysia (Borneo).
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  • A pair of the largest dung beetles in Borneo (Catharsius dayacus) clean up after the droppings of a Sambar Deer in the rainforest of Danum Valley, Sabah. Although this species does not roll balls like its more well-known relatives, it nevertheless is responsible for burying large amounts of animal scat and provides a vital ecological service. Nearly a hundred species of dung beetles have been recorded from Danum alone; studies have shown that their diversity not only directly reflects the wide assortment of vertebrates in an area but also can serve as a valuable indicator of habitat heterogeneity and forest disturbance. Sabah, Malaysia (Borneo).
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  • Dung Beetle (Epilissus splendidus). Mantadia National Park, Madagascar.
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  • Leaf of Colocasia gigantea with round holes left by feeding Chrysomelid beetle (Aplosonyx ancora). The beetle bites circular leaf trenches to drain the leaf of its toxic latex prior to consumption. Ha Giang, Vietnam.
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