Chien C. Lee

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  • Iridescent Bark Mantis (Metallyticus splendidus), juvenile. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • The enormous caterpillars of the Atlas Moth (Attacus atlas) consume huge amounts of leaves before they will pupate. Sarawak, Malaysia (Borneo).
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  • The hypodermic needle-like bristles on this slug moth caterpillar (Thosea sp., family Limacodidae) are capable of inflicting painful stings, making them an effective defense against would-be predators. Sarawak, Malaysia (Borneo).
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  • Spotted Cuscus (Spilocuscus maculatus chrysorrhous), juvenile. Central Papua, Indonesia (New Guinea).
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  • Common Mormon Swallowtail (Papilio polytes). This young butterfly caterpillar avoids predation by appearing unappealingly similar to a bird dropping. Sarawak, Malaysia (Borneo).
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  • Bagworm caterpillars (moths of the family Psychidae) are known for their habit of hiding themselves in a portable shelter of plant materials that they secure with silken threads, with each species having its own preference of design. This species begins with small accurately trimmed twigs formed in a 3-sided spiral pyramid, and adds additional larger twigs at the end as the caterpillar grows in size, in a masterful example of miniature craftsmanship. East Kalimantan, Indonesia (Borneo).
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  • A young Sunda Colugo (Galeopterus variegatus) peers out from the embrace of its mother's membranous wings. When fully grown it will be able to glide great distances from tree to tree in their nightly search for edible leaves, flowers, and sap. Also known as Flying Lemurs, they bear no relation to true lemurs, and in fact are so unique that they have been placed in their own mammalian order. Bako National Park, Sarawak, Malaysia (Borneo).
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  • Although each armed with stinging bristles, a brood of juvenile nettle caterpillars (Limacodidae) enhances their defense by forming a tight grouping. Each caterpillar faces outward, presenting a wall of venomous spines to their enemies. Sarawak, Malaysia (Borneo).
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  • Treehoppers (Alchisme inermis), adult with nymphs. Parque Natural Chicaque, Cundinamarca, Colombia.
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  • Malayan Lacewing Butterfly (Cethosia hypsea) caterpillars utilize warning coloration to signify to predators that they are unpalatable. Sarawak, Malaysia (Borneo).
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  • Unable to fly with a load many times her own weight, a female Thread-waisted Sand Wasp (Ammophila beniniensis) hauls a large caterpillar which she has paralyzed with her sting. She is seeking a burrow she has prepared in advance and, even though it may be a considerable distance away, she shows a remarkable orientation ability even though there may be numerous obstacles in her path. Once placed inside, with the burrow sealed shut by small rocks and its entrance hidden from potential thieves, the ill-fated caterpillar will serve as live food for the single egg the wasp has deposited along with it. Fianarantsoa, Madagascar.
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  • Sunda Colugo (Galeopterus variegatus), adult female with young. Sarawak, Malaysia (Borneo).
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  • A butterfly caterpillar (Cupha erymanthis) standing guard over a clutch of parasitic wasp pupae that recently hatched from its body. Having been impregnated by eggs from a female wasp, the larvae develop within the caterpillar, consuming its flesh as they grow. In order to keep their host alive, the wasp larvae are careful to avoid all vital organs and the caterpillar thus appears and behaves rather normal until the larvae eventually emerge. In a similar vein to the mind-controlled zombie ants, the wasp larvae also secrete some cocktail of hormones that modify the behavior of the caterpillar, in this case it is induced to stand guard and protect the wasp pupae from predators or other parasitic wasps until it eventually dies of starvation. Sarawak, Malaysia (Borneo).
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  • A male Rough Guardian Frog (Limnonectes finchi) transports tadpoles on its back to a suitable pool of water where it will release them. Sabah, Malaysia (Borneo).
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  • Shag-carpet caterpillar (Prothysana sp.). Sumaco Napo-Galeras National Park, Napo, Ecuador.
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  • Bagworm (family Psychidae) moth caterpillar. Ranomafana National Park, Madagascar.
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  • Bright colors warn potential predators of the painful stinging bristles worn by this slug moth caterpillar (family Limacodidae). Mulu National Park, Sarawak, Malaysia (Borneo).
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  • A juvenile Spotted Cuscus forages in a small tree for leaves and fruit at night. Central Papua, Indonesia (New Guinea).
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  • Green Tree Python (Morelia azurea), juvenile.
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  • Giant Leaf Katydid (Pseudophyllus hercules), juvenile female resting by day on foliage of a Phanera vine. Sarawak, Malaysia (Borneo).
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  • Banded Swallowtail (Papilio demolion), gregarious caterpillars. Sarawak, Malaysia (Borneo).
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  • After several months of feeding on Aristolochia leaves, the mature caterpillar of this large birdwing butterfly (Troides andromache) prepares for its final moult into the pupal stage when metamorphosis will transform it into an adult. Sabah, Malaysia (Borneo).
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  • Young Ring-tailed Lemurs (Lemur catta) will transition from riding on the belly to the back of their mother after 4-6 weeks of age. Amboasary, Madagascar.
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  • Mossy masquerade: a young Moss Mimic Katydid (Championica montana) doing what it does best. If you specialize in eating moss it’s a big plus if you look like it as well. In the damp cloud forests of Costa Rica, every surface is covered with dripping wet bryophytes, such as the leaf this katydid is perched on. Although abundant, it’s not the most energy rich food in the world, so these insects conserve their energy by moving slowly while they graze. Faced with a potential threat, they simply lay flat against the mossy substrate and disappear. Limón, Costa Rica.
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  • Almost indistinguishable from the branch on which it rests, a Papuan Frogmouth (Podargus papuensis) perches motionless next to its chick. Like other frogmouths, these birds scarcely build any nest at all and rely on camouflage to escape the detection of predators. Papua, Indonesia (New Guinea).
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  • Spiny katydid (Panacanthus spinosus), juvenile. Coclé, Panama.
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  • Katydid (Scambophyllum sp.), juvenile female. Sarawak, Malaysia (Borneo).
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  • Paradise Tree Snake (Chrysopelea paradisi). Sarawak, Malaysia (Borneo).
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  • A juvenile leaf insect (Nanophyllium australianum) from the rainforest of Iron Range National Park. Queensland, Australia.
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  • The erratic feeding pattern of this juvenile Malaysian Katydid (Ancylecha fenestrata) imparts necrotic scars on the foliage of its host plant (Glochidion rubrum). When combined with the insect's irregular green and brown coloration, this amplifies its camouflage while it rests on the underside of a leaf. Sabah, Malaysia (Borneo).
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  • Hooktip moth caterpillar (family Drepanidae). Sabah, Malaysia (Borneo)
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  • Marauder Ants (Carebara affinis) attacking a beetle grub. The minor workers are assisted by their larger sisters, the majors and super-majors, which use their powerful mandibles to cut the grub into smaller pieces. Sarawak, Malaysia (Borneo).
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  • Almost looking like it belongs more on a coral reef than in a rainforest, this moth caterpillar (Idonauton cf. apicalis) uses bright aposematic patterns to warn potential predators of its stinging bristles. Caterpillars of this family (Limacodidae) are often called ‘slug moths’ because they lack prolegs and instead move over the surface of a leaf in an oozing-fashion using suckers. Sabah, Malaysia (Borneo).
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  • Dragon-headed Katydid (Lesina blanchardi), juvenile. Sarawak, Malaysia (Borneo).
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  • Kirindy Leaf-toed Gecko (Paroedura rennerae), juvenile. Tsingy de Bemaraha National Park, Madagascar.
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  • Spiny helmeted katydid (Sasima sp.) from the rainforest of southern New Guinea. The bristly armaments on this juvenile specimen serve not only to deter predators, but also to disguise the insect in the mossy forest understory. Helmet katydids (Phyllophorinae) comprise some of the world's largest species. Digul River, South Papua, Indonesia (New Guinea).
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  • Sunda Pangolin (Manis javanica), juvenile. Sarawak, Malaysia (Borneo).
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  • Like other langurs, Silvered Leaf Monkeys (Trachypithecus cristatus) are highly social and travel in cohesive groups numbering up to 40 or more individuals. They have a specialized diet, feeding almost exclusively on high-protein leaves. Sarawak, Malaysia (Borneo).
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  • A juvenile Parson's Chameleon (Calumma parsoni). More frequently occurring in shades of green, this is one of the world's largest chameleons, with adult specimens sometimes reaching the size of a house cat. They are endemic to rainforest habitats on Madagascar and are threatened by both habitat loss and collection for the international pet trade. Andasibe, Madagascar.
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  • Under the cover of darkness, a cicada sheds its nymphal skin to emerge as a fully grown adult.  Sarawak, Malaysia (Borneo).
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  • A juvenile leaf insect (Nanophyllium australianum) from the rainforest of Iron Range National Park. Queensland, Australia.
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  • With an adult size of less than one meter in length, the beautifully patterned Children's Python (Antaresia childreni) is one of the smallest pythons. Cape Range National Park, Exmouth, Western Australia.
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  • The endangered Bear Monkey (Trachypithecus vetulus monticola) is endemic the mountain forests of Sri Lanka. Central Province, Sri Lanka.
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  • Leopard (Panthera pardus), juvenile. Ngamiland, Botswana.
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  • A juvenile Rock Monitor (Varanus albigularis). Mpumalanga, South Africa.
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  • Elibia dolichus, caterpillar. Although large and seemingly intimidating, this sphinx moth caterpillar is completely harmless and relies entirely on camouflage and eyespots to deter predators. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Marauder Ants (Carebara affinis) attacking a beetle grub. The minor workers are assisted by their larger sisters, the majors and super-majors, which use their powerful mandibles to cut the grub into smaller pieces. Sarawak, Malaysia (Borneo).
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  • Bushy-crested Hornbill (Anorrhinus galeritus), juvenile. Sepilok Forest Reserve, Sabah, Malaysia (Borneo).
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  • In the coastal forests of northern Borneo, a young Silvery Lutung (Trachypithecus cristatus) is groomed by its mother. Like many leaf monkey species, these highly social primates display brightly colored fur when born that gradually fades after several months into their adult coloration. The exact reason for this adaption is uncertain, but it may help in protection: members of the group can easily distinguish their vulnerable young at a glance, or locate them quickly in the dense foliage of the rainforest canopy. Another interesting theory is that certain predators such as Clouded Leopards, are red/green colorblind, so an orangish pelt may actually be an effective camouflage. Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Almost looking like it belongs more on a coral reef than in a rainforest, this moth caterpillar (Idonauton cf. apicalis) uses bright aposematic patterns to warn potential predators of its stinging bristles. Caterpillars of this family (Limacodidae) are often called ‘slug moths’ because they lack prolegs and instead move over the surface of a leaf in an oozing-fashion using suckers. Sabah, Malaysia (Borneo).
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  • The bright orange coloration of this infant Silvered Langur (Trachypithecus cristatus) helps them to be seen and hence guarded by the entire troop. They will gradually lose this coloration after 3 months of age. Bako National Park, Sarawak, Malaysia (Borneo).
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  • Paradise Tree Snake (Chrysopelea paradisi). Sarawak, Malaysia (Borneo).
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  • Just a few millimeters in length, a tiny froglet (Microhyla nepenthicola) makes its first climb out of the watery chamber of the pitcher plant in which it was born. This species is exclusively reliant on pitcher plants for depositing their eggs. Sarawak, Malaysia (Borneo).
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  • Deep within a cave in Mulu National Park, the sticky threads secreted by the larvae of a fungus midge (family Mycetophilidae) function as a trap for small flying insects.  When ensnared in this trap, prey are reeled in and eaten. Sarawak, Malaysia (Borneo).
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  • Within the microcosm of a mossy tree stump, a tiny hidden predator lies patiently in wait. Still a juvenile, this praying mantis (Haania sp.) will eventually mature and shed most of its moss-like camouflage in favor of fully developed wings. Sarawak, Malaysia (Borneo).
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  • A newly hatched Green Turtle (Chelonia mydas) makes its way down the beach and feels the touch of the sea for the first time in its life. Many predators lie in wait and young sea turtles have an astonishingly low survival rate, but if it reaches adulthood it may well return to the same beach years in the future to lay eggs of its own. Sarawak, Malaysia (Borneo).
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  • Newly hatched Red Stink Bugs (Pycanum alternatum) cluster together to amplify the effect of their aposematic warning coloration. Sarawak, Malaysia (Borneo).
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  • Like all cobras, this juvenile Equatorial Spitting Cobra (Naja sumatrana), also called the Black Spitting Cobra and Sumatran Spitting Cobra, inflates its hood when approached by a potential threat. As an additional defense this species is also capable of spitting venom at distances of up to 1 meter, which is typically aimed with great accuracy at the eyes of its adversary. Sarawak, Malaysia (Borneo).
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  • Venezuela’s fabled tabletop mountains may have once inspired Arthur Conan Doyle’s science fiction novel “The Lost World”, where he imagined the summits to be home to a plethora of gigantic prehistoric creatures that had been separated from the evolution of life on the rest of the planet for eons of time. Disappointingly, dinosaurs were never found on these mountains and, despite being home to many endemics and indeed a fair share of “living fossils”, recent genetic studies have shown that many species in fact have likely arrived well after the tepuis had been fully formed. This reveals that the imposing rocky cliffs, which seemingly isolate the summits from the jungles below, may not be as significant a physical barrier for dispersal as once believed. This young Boddaert's Tropical Racer (Mastigodryas boddaerti), encountered on the plateau of Auyán Tepui, is a widespread species in northern South America, and although prey is scarce on these mountaintops, it presumably hunts for endemic frogs.  Canaima National Park, Venezuela.
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  • Moth caterpillar (Synchlora sp.), disguised with plant detritus. Soberanía National Park, Panama.
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  • Canopy Slug-eating Snake (Sibon canopy), a new species described in January 2023, endemic to Panama. Panamá Oeste, Panama.
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  • Verreaux's Sifaka (Propithecus verreauxi), mother with young. Amboasary, Madagascar.
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  • Ant-mimicking mantis (Hapalopeza sp.), juvenile. Sarawak, Malaysia (Borneo)
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  • This unidentified caterpillar which feeds on epiphytic moss, is camouflaged to look like its preferred substrate. Arfak Mountains, West Papua, Indonesia (New Guinea).
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  • Babbling Torrenteer (Hyloscirtus alytolylax), young froglet metamorph. Mindo, Ecuador.
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  • Sunda Pangolin (Manis javanica), juvenile. Sarawak, Malaysia (Borneo).
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  • A female Bornean Orang-utan (Pongo pygmaeus) with her young infant. Semengoh Forest Reserve, Sarawak, Malaysia (Borneo).
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  • Smooth Guardian Frog (Limnonectes palavanensis), male transporting tadpoles on back. Sarawak, Malaysia (Borneo).
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  • Planthopper (Flatidae), nymph. Ranomafana National Park, Madagascar.
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  • A column of nomadic Asian army ants (Aenictus laeviceps) carry their developing larvae to a new temporary nesting site. Ants of the genus Aenictus are specialized predators on other social insects, including wasps, bees, termites, and other ant species. Sarawak, Malaysia (Borneo).
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