Chien C. Lee

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  • The karst of the Melinau Limestone Formation gives Mulu National Park its rugged terrain. These forests and caves abound with many endemic species. Sarawak, Malaysia (Borneo).
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  • The steep karst formations of the Subis Limestone Complex of Niah National Park hold numerous large caverns, among which are numerous important archaeological sites. The presence of humans in Niah Cave has been dated to about 40,000 years ago, making it the oldest known settlement in East Malaysia. Sarawak, Malaysia (Borneo).
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  • Uplifted by the collision of continents, the jagged limestone peaks of New Guinea’s Central Range rise to nearly 5000 meters in height. The alpine vegetation of these remote mountain ridges and valleys are home to a rich assortment of endemic plants and animals, many of which are new to science. Highland Papua, Indonesia (New Guinea).
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  • Uplifted by the collision of continents, the jagged limestone peaks of New Guinea’s Central Range rise to nearly 5000 meters in height. Here, at the edge of the island’s divide, thick clouds rise from the rainforest plains of the south and push up against towering cliffs. Papua, Indonesia.
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  • Uplifted by the collision of continents, the jagged limestone peaks of New Guinea’s Central Range rise to nearly 5000 meters in height. The alpine vegetation of these remote mountain ridges and valleys are home to a rich assortment of endemic plants and animals, many of which are new to science. Highland Papua, Indonesia (New Guinea).
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  • The spectacular karst complex of the Sangkulirang Peninsula is the most extensive limestone area in Borneo.  This region is poorly explored but is believed to harbour a rich diversity of endemic plants. East Kalimantan, Indonesia.
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  • The limestone mountains of Mulu National Park are some of the highest and most spectacular in all Southeast Asia, having been uplifted and eroded by weather over millions of years. The famous 'Pinnacles' occur on the shoulder of Gunung Api, with the prominent Gunung Benarat seen in the background. Sarawak, Malaysia (Borneo).
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  • Weathering from rain and wind has given rise to these razor-sharp limestone pinnacles, which can be found on the upper slopes of Mount Api in Mulu National Park. Sarawak, Malaysia (Borneo).
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  • Forest on limestone hill. Ha Giang, Vietnam.
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  • A crystal-clear stream flows through the forest at Khao Nor Chuchi Wildlife Sanctuary.  These spring-fed waters which emerge from limestone rock are highly alkaline and contain high amounts of dissolved calcium carbonate. Krabi, Thailand.
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  • The incredible labyrinth of limestone islands within Waigeo's Kabui Bay. Lying on the northern edge of Raja Ampat, this region is home to the world's richest marine life, including 1500 species of fish and 75% of all known coral species. The islands themselves also host a diverse assortment of rainforest flora and fauna, including slipper orchids, endemic marsupials, and birds of paradise. West Papua, Indonesia.
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  • Stone forest at dawn: the first light of day brushes the limestone pinnacles of Madagascar’s Tsingy de Bemaraha National Park. Spanning 600-square miles, this vast and impenetrable labyrinth of knife-edged towers and slot canyons is an invaluable wilderness reserve; 90% of all species occurring here are found nowhere else on Earth.
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  • A pair of Lesser False Vampire Bats (Megaderma spasma) roosting near the entrance of a shallow limestone cave. Named because of their superficial resemblance to true vampire bats (which are restricted to Central and South America), Megaderma are insectivorous and do not drink blood. This species sometimes attains a large size (nearly 35g in weight) and is known to occasionally take more substantial prey including lizards, small birds, mammals, and even other bats. Sarawak, Malaysia (Borneo).
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  • Sunlight filters through lowland rainforest into the entrance of a limestone cave in Mulu National Park. Sarawak, Malaysia (Borneo).
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  • Cave crab (Stygothelphusa cranbrooki), endemic to a single limestone cave in western Borneo. Like other cave-adapted crabs it has elongated legs and has lost most of its pigment. However, although living in complete darkness it has not yet lost function of its eyes. Sarawak, Malaysia (Borneo).
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  • With its varied topography of limestone mountains and lowland rainforest, Mulu National Park is home to one of the richest floral diversities in Borneo. Sarawak, Malaysia (Borneo).
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  • Summit ridge of Gunung Api, the highest limestone mountain in Borneo. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Pitcher plant (Nepenthes campanulata), a limestone lithophytic carnivorous plant, here growing on a cliff 250m above the rainforest canopy. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • A Lesser False Vampire Bat (Megaderma spasma) roosting near the entrance of a shallow limestone cave. Named because of their superficial resemblance to true vampire bats (which are restricted to Central and South America), Megaderma are insectivorous and do not drink blood. This species sometimes attains a large size (nearly 35g in weight) and is known to occasionally take more substantial prey including lizards, small birds, mammals, and even other bats. Sarawak, Malaysia (Borneo).
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  • The Beauty Ratsnake (Elaphe taeniura grabowskyi) is an adept climber, capable of scaling limestone cave walls where it feeds on roosting bats and swiftlets. Sarawak, Malaysia (Borneo).
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  • Helmet Orchid (Corybas crenulatus). This species is endemic to Borneo, where it grows in thick moss on limestone rocks. Sarawak, Malaysia (Borneo).
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  • Perched high on a limestone cliff, a rare slipper orchid (Paphiopedilum stonei) puts on a showy display of multiple blooms. This endangered species occurs only in a few localities in western Borneo where its population has been heavily depleted by wild collecting. Sarawak, Malaysia (Borneo).
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  • A non-climbing species, this pitcher plant (Nepenthes campanulata) grows only on the vertical faces of tall limestone cliffs in Borneo. It is so difficult to observe in habitat that it was once thought to be extinct. Sarawak, Malaysia (Borneo).
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  • Helmet Orchid (Corybas crenulatus). This species is endemic to Borneo, where it grows in thick moss on limestone rocks. Sarawak, Malaysia (Borneo).
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  • Compact clumps of the rare pitcher plant Nepenthes campanulata grow on the vertical walls of a limestone cliff nearly a hundred meters above the canopy of the rainforest. Sarawak, Malaysia (Borneo).
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  • A female cave cricket (Diestrammena sp.) pushes her ovipositor into the spongy surface of the limestone cave wall where she will lay her eggs. Sarawak, Malaysia (Borneo).
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  • Round-eyed Gecko (Cnemaspis sp. aff. paripari), undescribed and one of several species of Cnemaspis which are endemic to separate and isolated limestone outrops. Sarawak, Malaysia (Borneo).
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  • Impressive trees of this enormous Fishtail Palm grow at the entrance of a limestone cave in Mulu National Park. These and many of the other trees growing in the vicinity have grown from seeds carried there by fruit-eating bats which roost in the cave. Sarawak, Malaysia (Borneo).
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  • A pair of Lesser False Vampire Bats (Megaderma spasma) roosting near the entrance of a shallow limestone cave. Named because of their superficial resemblance to true vampire bats (which are restricted to Central and South America), Megaderma are insectivorous and do not drink blood. This species sometimes attains a large size (nearly 35g in weight) and is known to occasionally take more substantial prey including lizards, small birds, mammals, and even other bats. Sarawak, Malaysia (Borneo).
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  • Nepenthes mapuluensis, a rare pitcher plant endemic to the limestone mountains of East Kalimantan, Indonesia (Borneo).
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  • Pitcher plant (Nepenthes northiana). Sarawak, Malaysia (Borneo).
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  • Begonia pulvinifera. Ha Giang, Vietnam.
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  • Niah Cave Gecko (Cyrtodactylus cavernicolus). Sarawak, Malaysia (Borneo).
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  • A clearing in raiforest is dominated by wild banana trees (Musa sp.). Ha Giang, Vietnam.
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  • Pitcher plant (Nepenthes campanulata). Sarawak, Malaysia (Borneo).
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  • Cave cricket (Diestrammena sp.). Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Deer Cave (located in Mulu National Park) is one of the world's largest cave passages, with an opening chamber over 120 meters in height. It is home to an estimated 5 million bats. Sarawak, Malaysia (Borneo).
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  • Fairy lantern (Thismia ornata). Sarawak, Malaysia (Borneo).
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  • Deep within a cave in Mulu National Park, the sticky threads secreted by the larvae of a fungus midge (family Mycetophilidae) function as a trap for small flying insects.  When ensnared in this trap, prey are reeled in and eaten. Sarawak, Malaysia (Borneo).
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  • Eerie smokeless fires dance among fissures in the limestone rocks where ignited natural gas escapes in Tanjung Api (Fire Cape) National Park. Central Sulawesi, Indonesia.
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  • Land snail (Vitrinula moluensis), a rare species endemic to limestone in Sarawak. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Closeup of the colorful inflorescence of Amorphophallus julaihii. Like other aroids, the small male flower (visible) are located above the female flowers (hidden inside the spathe). This species is endemic to limestone forest in northern Borneo. Sarawak, Malaysia.
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