Chien C. Lee

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  • Proboscis Monkey (Nasalis larvatus). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • One of Borneo's most famous endemic mammals, the Proboscis Monkey (Nasalis larvatus) is restricted to coastal swamp forests and mangroves. Bako National Park, Sarawak, Malaysia (Borneo).
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  • Proboscis Monkey (Nasalis larvatus), male. Sarawak, Malaysia (Borneo).
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  • Proboscis Monkey (Nasalis larvatus), subadult male. Sarawak, Malaysia (Borneo).
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  • An adult male Silvered Leaf Monkey (Trachypithecus cristatus) keeps a watchful eye out for any threats to his harem of females. Sarawak, Malaysia (Borneo).
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  • Silvered Leaf Monkey (Trachypithecus cristatus). Sarawak, Malaysia (Borneo).
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  • The endangered Bear Monkey (Trachypithecus vetulus monticola) is endemic the mountain forests of Sri Lanka. Central Province, Sri Lanka.
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  • Silvered Leaf Monkey (Trachypithecus cristatus), rare white morph. Bako National Park, Sarawak, Malaysia (Borneo).
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  • Silvered Leaf Monkey (Trachypithecus cristatus). Sarawak, Malaysia (Borneo).
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  • Like other langurs, Silvered Leaf Monkeys (Trachypithecus cristatus) are highly social and travel in cohesive groups numbering up to 40 or more individuals. They have a specialized diet, feeding almost exclusively on high-protein leaves. Sarawak, Malaysia (Borneo).
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  • In the coastal forests of northern Borneo, a young Silvery Lutung (Trachypithecus cristatus) is groomed by its mother. Like many leaf monkey species, these highly social primates display brightly colored fur when born that gradually fades after several months into their adult coloration. The exact reason for this adaption is uncertain, but it may help in protection: members of the group can easily distinguish their vulnerable young at a glance, or locate them quickly in the dense foliage of the rainforest canopy. Another interesting theory is that certain predators such as Clouded Leopards, are red/green colorblind, so an orangish pelt may actually be an effective camouflage. Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • A large group of Proboscis Monkeys (Nasalis larvatus) prepares for sleep in a tree overhanging a river. Sabah, Malaysia (Borneo).
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  • The Maroon Langur (Presbytis rubicunda), also known as the Red Leaf Monkey, is endemic to Borneo and nearby Karimata Island, with several subspecies occurring throughout its range. Like other leaf monkeys they are almost completely arboreal and feed on foliage, seeds, and some fruits. Sabah, Malaysia.
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  • The Crested Black Macaque (Macaca nigra nigra) is one of four monkey species endemic to Sulawesi.  They are sometimes wrongly referred to as apes because their tail is extremely small and difficult to see.  They travel mostly on the ground in large groups of up to nearly 100 individuals. North Sulawesi, Indonesia.
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  • Threatened with encroaching cultivation and forest loss, a Southern Purple-Faced Langur (Trachypithecus vetulus vetulus) enters a tea plantation to feed on the fresh young leaves. These endangered monkeys are endemic to Sri Lanka’s southern wet forests, and although once widespread, are now facing severe habitat fragmentation due to roads and settlements. This can lead to human conflicts where the monkeys are inevitably forced to exploit agricultural areas for food. Sinharaja National Park. Sri Lanka.
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  • A Red-shanked Douc (Pygathrix nemaeus) munches on a handful of young vine leaves it has picked in the forest canopy. Although these endangered monkeys feed on over sixty different species of plants, they select their food carefully and in particular avoid sweet fruits which can upset the balance of bacteria in their guts. Like other leaf-eating colobine monkeys, doucs have large chambered stomachs – making them the only ruminant primates and giving them their characteristic pot-bellied appearance. Son Tra Nature Reserve, Vietnam.
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  • A Maroon Langur (Presbytis rubicunda) pauses for a break while munching on his chosen leaf. Endemic to Borneo and a few offshore islands, these leaf monkeys feed on a wide range of foliage and unripe fruits, traveling in groups of up to a dozen individuals. One theory regarding their distinctive bright orange coloration is that this may help to confuse some predators into mistaking them for a much larger and stronger Orangutan, and thus deter attacks. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • The Maroon Langur (Presbytis rubicunda) is endemic to Borneo and nearby Karimata Island, with several subspecies occurring throughout its range. Like other leaf monkeys they are almost completely arboreal and feed on foliage, seeds, and some fruits.  Danum Valley Conservation Area, Sabah, Malaysia.
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  • The Maroon Langur (Presbytis rubicunda) is endemic to Borneo and nearby Karimata Island, with several subspecies occurring throughout its range. Like other leaf monkeys they are almost completely arboreal and feed on foliage, seeds, and some fruits. Sabah, Malaysia.
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  • The Sumatran Surili (Presbytis melalophos) is endemic to the island of Sumatra.  Like other leaf monkeys, this species is primarily folivorous, but also consumes flowers and fruits. Kerinci Seblat National Park, West Sumatra, Indonesia.
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  • The least known of all of Sulawesi's monkeys, Heck's Macaque (Macaca hecki) occurs on the northwestern portion of the island. Five other endemic species of macaque are distributed in different regions of Sulawesi, occasionally producing hybrids where their ranges overlap. All are diurnal and feed largely on fruits. Central Sulawesi, Indonesia.
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  • The least known of all of Sulawesi's monkeys, Heck's Macaque occurs on the northwestern portion of the island. Five other endemic species of macaque are distributed in different regions of Sulawesi, occasionally producing hybrids where their ranges overlap. All are diurnal and feed largely on fruits. Central Sulawesi, Indonesia.
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  • The least known of all of Sulawesi's monkeys, Heck's Macaque occurs on the northwestern portion of the island. Five other endemic species of macaque are distributed in different regions of Sulawesi, occasionally producing hybrids where their ranges overlap. All are diurnal and feed largely on fruits. Central Sulawesi, Indonesia.
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  • Sri Lankan Grey Langur (Semnopithecus priam thersites). Yala National Park, Sri Lanka.
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  • The bright orange coloration of this infant Silvered Langur (Trachypithecus cristatus) helps them to be seen and hence guarded by the entire troop. They will gradually lose this coloration after 3 months of age. Bako National Park, Sarawak, Malaysia (Borneo).
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  • Southern Purple-faced Langur (Trachypithecus vetulus vetulus). Sinharaja National Park, Sri Lanka.
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  • Maroon Langur (Presbytis rubicunda) feeding on leaves of Mikania micrantha, an invasive vine. Danum valley, Sabah, Malaysia (Borneo).
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  • One of the world's most endangered primates, the Bornean Banded Langur (Presbytis chrysomelas chrysomelas) has been threatened by both habitat loss and hunting. Now, only a few isolated populations remain in coastal Sarawak, with an estimated total of just a few hundred individuals. They feed on foliage and fruits in lowland rainforest. Sarawak, Malaysia (Borneo).
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  • Threatened by habitat loss and hunting pressure, the Crested Black Macaque (Macaca nigra nigra) is now listed as Critically Endangered. This species lives only in forested areas at the northernmost tip of the island of Sulawesi. North Sulawesi, Indonesia.
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  • Marañón White-fronted Capuchin (Cebus yuracus). Yasuní National Park, Ecuador.
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  • Napo Saki (Pithecia napensis). Yasuní National Park, Ecuador.
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  • Geoffroy’s Tamarin (Saguinus geoffroyi). Colón, Panama.
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  • With their vertical pupils and opposable thumbs, Phyllomedusa are easily some of the most distinguishable of all tree frogs. However, it is their peculiar form of locomotion that is perhaps their most striking feature: when seen walking slowly down a branch with their bodies raised, they appear decidedly ‘un-froglike’. Studies into their anatomy reveal that they possess a specialized musculature that not only allows them to move their limbs independently (as opposed to the simultaneous bilateral hopping motions of most other frogs), but also gives them a ‘precision grip’ which enables steady movement over thin branches. This grasping ability is rare among animals and is shared predominantly with higher primates, hence leading to one of their common names: ‘Monkey Frogs’. Coca, Ecuador.
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  • Pallid Monkey Moth (Sphingognatha asclepiades). Sarawak, Malaysia (Borneo).
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  • Tarsier Monkey-Frog (Phyllomedusa tarsius). Yasuní National Park, Ecuador.
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  • This small dung beetle (Paragymnopleurus maurus) is preparing a ball made from monkey droppings in which it will deposit its eggs and then bury. Sarawak, Malaysia (Borneo).
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  • Morning sunlight touches the forested crest of the continental divide in El Triunfo Biosphere Reserve, southern Mexico. Spanning nearly 120,000 hectares in the Sierra Madre de Chiapas, this reserve protects vital ecosystems in one of the most biodiverse parts of the country; it is one of the few places in Mexico where rare species such as the Resplendent Quetzal, Horned Guan, and Geoffroy’s Spider Monkey can be found. Although protected since 1990, the area nevertheless continues to be threatened by uncontrolled fires, road development, and landslides which are caused by forest clearance.
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  • A large male Bornean Clouded Leopard (Neofelis diardi) is caught on camera trap as he patrols his territory in the rainforest of Danum Valley, Sabah. Borneo's largest cat species, the clouded leopard preys on a variety of smaller wildlife ranging from deer and wild boar to monkeys. Sabah, Malaysia (Borneo).
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  • Bornean Clouded Leopard (Neofelis diardi borneensis), captive female. This is the largest land predator in Borneo and possesses the longest canine teeth of any living feline. It is a nocturnal hunter and has been observed feeding on prey as large as large as Proboscis Monkeys. Recently this species was distinguished from the mainland Clouded Leopard (Neofelis nebulosa). Sarawak, Malaysia (Borneo).
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  • One of the world’s least studied crocodilians, the False Gharial (Tomistoma schlegelii) is a rare inhabitant of the dark tannin colored waters of peat swamps in Borneo, Sumatra, and Malaysia. Due to their narrow snout they were once believed to feed exclusively on fish, however large specimens can reach lengths of over 5 meters and have more recently been observed to consume prey as large as Proboscis Monkeys, or in one unfortunate instance, a grown man. Although much remains unknown about their life history, it is clear that they are rapidly becoming endangered due to destruction of their peat swamp habitats. Sarawak, Malaysia (Borneo).
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