Chien C. Lee

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  • A Pygmy Loris (Nycticebus pygmaeus) makes a meal of a giant stick insect (Tirachoidea sp.).  Although primarily insectivorous, these nocturnal primates will also feed on fruit, sap, flowers, lizards, and nestling birds.  Captive. Endangered Primate Rescue Centre, Cuc Phuong National Park, Ninh Binh, Vietnam.
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  • The Western Tarsier (Cephalopachus bancanus borneanus), also known as Horsfield's Tarsier, is a small nocturnal primate occurring in the lowland forests of Sumatra and Borneo. They are carnivorous and feed mainly on insects, but occasionally take birds and bats. Sarawak, Malaysia (Borneo).
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  • Nycticebus menagensis. One of four species of Slow Loris found in Borneo. All Slow Loris are nocturnal arboreal primates which feed on small animals, insects and soft fruits. Sarawak, Malaysia (Borneo).
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  • Feared by some, revered by others, the Aye-aye (Daubentonia madagascariensis) is surely one of Madagascar's most bizarre lemurs. Unique among primates, the Aye-aye's front incisors grow continuously like a rodent, leading to the early belief that they were some sort of gigantic nocturnal squirrel. Equally unusual are its long bony middle fingers which are used by the animal to tap tree trunks for the sound of insect grubs inside and then extract them like a fish hook. Unfortunately, Aye-ayes have become gravely endangered from habitat loss and persecution by people: they are often killed on sight due to the superstition that they are an omen of death. Tomasina, Madagascar.
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  • A Gursky's Spectral Tarsier (Tarsius spectrumgurskyae) perches outside of its tree hole roost. These tiny nocturnal primates emerge every evening to hunt for insects. Tangkoko Nature Reserve, North Sulawesi, Indonesia.
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  • In the coastal forests of northern Borneo, a young Silvery Lutung (Trachypithecus cristatus) is groomed by its mother. Like many leaf monkey species, these highly social primates display brightly colored fur when born that gradually fades after several months into their adult coloration. The exact reason for this adaption is uncertain, but it may help in protection: members of the group can easily distinguish their vulnerable young at a glance, or locate them quickly in the dense foliage of the rainforest canopy. Another interesting theory is that certain predators such as Clouded Leopards, are red/green colorblind, so an orangish pelt may actually be an effective camouflage. Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • A Red-shanked Douc (Pygathrix nemaeus) munches on a handful of young vine leaves it has picked in the forest canopy. Although these endangered monkeys feed on over sixty different species of plants, they select their food carefully and in particular avoid sweet fruits which can upset the balance of bacteria in their guts. Like other leaf-eating colobine monkeys, doucs have large chambered stomachs – making them the only ruminant primates and giving them their characteristic pot-bellied appearance. Son Tra Nature Reserve, Vietnam.
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  • A Gursky's Spectral Tarsier (Tarsius spectrumgurskyae) perches outside of its tree hole roost. These tiny nocturnal primates emerge every evening to hunt for insects. Tangkoko Nature Reserve, North Sulawesi, Indonesia.
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  • One of the world's most endangered primates, the Bornean Banded Langur (Presbytis chrysomelas chrysomelas) has been threatened by both habitat loss and hunting. Now, only a few isolated populations remain in coastal Sarawak, with an estimated total of just a few hundred individuals. They feed on foliage and fruits in lowland rainforest. Sarawak, Malaysia (Borneo).
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  • A Horsfield's Tarsier (Cephalopachus bancanus) on the prowl for food in the rainforest of western Borneo. These tiny primates are exclusively carnivorous, hunting insects, small lizards, and even snakes by leaping deftly through the thick forest understory and snatching prey with their long nimble fingers. Unlike most other nocturnal mammals which possess a reflective layer behind their retina to help with night vision, tarsiers instead compensate by having enormous eyes. In fact, this species sports the largest eyes in relation to its body size of any mammal in the world - these are locked in place and the animal is only able to shift its vision by rotating its head. Sarawak, Malaysia (Borneo).
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  • Sri Lankan Grey Langur (Semnopithecus priam thersites). Yala National Park, Sri Lanka.
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  • Threatened by habitat loss and hunting pressure, the Crested Black Macaque (Macaca nigra nigra) is now listed as Critically Endangered. This species lives only in forested areas at the northernmost tip of the island of Sulawesi. North Sulawesi, Indonesia.
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  • Marañón White-fronted Capuchin (Cebus yuracus). Yasuní National Park, Ecuador.
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  • One of Borneo's most famous endemic mammals, the Proboscis Monkey (Nasalis larvatus) is restricted to coastal swamp forests and mangroves. Bako National Park, Sarawak, Malaysia (Borneo).
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  • The least known of all of Sulawesi's monkeys, Heck's Macaque (Macaca hecki) occurs on the northwestern portion of the island. Five other endemic species of macaque are distributed in different regions of Sulawesi, occasionally producing hybrids where their ranges overlap. All are diurnal and feed largely on fruits. Central Sulawesi, Indonesia.
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  • With its large stiff flower bracts and copious amounts of sucrose-laded nectar, Madagascar’s Traveler’s Palms (Ravenala spp.) have evolved to be pollinated almost exclusively by several species of lemurs. Here, a Common Brown Lemur (Eulemur fulvus) feeds at the infloresence of Ravenala hladikorum. Andasibe, Madagascar.
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  • The least known of all of Sulawesi's monkeys, Heck's Macaque occurs on the northwestern portion of the island. Five other endemic species of macaque are distributed in different regions of Sulawesi, occasionally producing hybrids where their ranges overlap. All are diurnal and feed largely on fruits. Central Sulawesi, Indonesia.
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  • Gursky's Spectral Tarsier (Tarsius spectrumgurskyae), juvenile. North Sulawesi, Indonesia.
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  • Goodman's Mouse Lemur (Microcebus lehilahytsara), feeding on the nectar of night-blooming Dombeya flowers. Andasibe, Madagascar.
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  • A Small-toothed Sportive Lemur (Lepilemur microdon) peers curiously from its tree hole in the montane forest of Ranomafana National Park, Madagascar. Usually emerging only at night, sportive lemurs frequently wake to keep a watchful eye out for intruders into their territory. Not long ago only 8 species of Lepilemur were known from Madagascar, but recent molecular work has resulted in at least 26 distinct species now being recognized, with more sure to follow.
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  • Bornean Orangutan (Pongo pygmaeus), infant. Sarawak, Malaysia (Borneo).
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  • The critically endangered Greater Bamboo Lemur (Prolemur simus) is restricted to wet rainforests of Madagascar's southeast where it is highly dependent on Cathariostachys bamboo. Ranomafana National Park, Madagascar.
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  • With fewer than a thousand individuals remaining in the wild, the Critically Endangered Golden Bamboo Lemur (Hapalemur aureus) holds on to a tenuous existence in the rainforests of Madagascar. It’s discovery in the mid-1980’s was a crucial factor that led to the foundation of the now famous Ranomafana National Park – a reserve that protects over 400 sq. km. of tropical forest. Like other bamboo lemurs, it feeds primarily on bamboo shoots which, due to their high cyanide content, are lethally toxic to most other animals. Ranomafana National Park, Madagascar.
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  • Silvered Leaf Monkey (Trachypithecus cristatus). Sarawak, Malaysia (Borneo).
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  • An adult male Silvered Leaf Monkey (Trachypithecus cristatus) keeps a watchful eye out for any threats to his harem of females. Sarawak, Malaysia (Borneo).
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  • Silvered Leaf Monkey (Trachypithecus cristatus), rare white morph. Bako National Park, Sarawak, Malaysia (Borneo).
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  • Maroon Langur (Presbytis rubicunda) feeding on leaves of Mikania micrantha, an invasive vine. Danum valley, Sabah, Malaysia (Borneo).
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  • Napo Saki (Pithecia napensis). Yasuní National Park, Ecuador.
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  • With fewer than a thousand individuals remaining in the wild, the Critically Endangered Golden Bamboo Lemur (Hapalemur aureus) holds on to a tenuous existence in the rainforests of Madagascar. It’s discovery in the mid-1980’s was a crucial factor that led to the foundation of the now famous Ranomafana National Park – a reserve that protects over 400 sq. km. of tropical forest. Like other bamboo lemurs, it feeds primarily on bamboo shoots which, due to their high cyanide content, are lethally toxic to most other animals. Ranomafana National Park, Madagascar.
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  • Ring-tailed Lemur (Lemur catta), sunbathing to warm themselves in the early morning. Berenty Private Reserve, Madagascar.
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  • The least known of all of Sulawesi's monkeys, Heck's Macaque occurs on the northwestern portion of the island. Five other endemic species of macaque are distributed in different regions of Sulawesi, occasionally producing hybrids where their ranges overlap. All are diurnal and feed largely on fruits. Central Sulawesi, Indonesia.
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  • The Maroon Langur (Presbytis rubicunda) is endemic to Borneo and nearby Karimata Island, with several subspecies occurring throughout its range. Like other leaf monkeys they are almost completely arboreal and feed on foliage, seeds, and some fruits.  Danum Valley Conservation Area, Sabah, Malaysia.
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  • A newly distinguished species, the Kayan Loris (Nycticebus kayan) differs from other Bornean loris species by a combination of morphological traits. Sarawak, Malaysia (Borneo).
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  • Although sometimes weighing over 9 kg, Indri (Indri indri) can make impressive leaps from tree to tree with their powerful long hind legs. Largest of the living lemurs, this critically-endangered species is an icon for the conservation of Madagascar's beleaguered wildlife, and its wailing song never fails to enchant visitors to their forest realm. Occuring only in the tropical rainforests on the island's eastern side, the Indri's natural range has dwindled due to widespread deforestation and hunting, despite their traditionally sacred status. Tomasina, Madagascar.
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  • A female Bornean Orang-utan (Pongo pygmaeus) with her young infant. Semengoh Forest Reserve, Sarawak, Malaysia (Borneo).
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  • Red-fronted Brown Lemur (Eulemur rufifrons)
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  • The endangered Bear Monkey (Trachypithecus vetulus monticola) is endemic the mountain forests of Sri Lanka. Central Province, Sri Lanka.
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  • Mountain Gorilla (Gorilla beringei beringei). Volcanoes National Park, Ruhengeri, Rwanda.
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  • With an adult size usually under 405g, the Pygmy Slow Loris (Nycticebus pygmaeus) is the smallest of all lorises. It is strictly nocturnal and survives on a diet of insects and fruit. Cuc Phuong National Park, Vietnam.
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  • Kayan Loris (Nycticebus kayan), juvenile. Gunung Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Milne-Edwards' Sifaka (Propithecus edwardsi). Ranomafana National Park, Madagascar.
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  • Northeast Bornean Orangutan (Pongo pygmaeus morio), adult male. Sabah, Malaysia (Borneo).
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  • Grey Mouse Lemur (Microcebus murinus). Kirindy Reserve, Morondava, Madagascar.
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  • Grey Mouse Lemur (Microcebus murinus). Kirindy Reserve, Madagascar.
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  • The Crested Black Macaque (Macaca nigra nigra) is one of four monkey species endemic to Sulawesi.  They are sometimes wrongly referred to as apes because their tail is extremely small and difficult to see.  They travel mostly on the ground in large groups of up to nearly 100 individuals. North Sulawesi, Indonesia.
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  • Proboscis Monkey (Nasalis larvatus), subadult male. Sarawak, Malaysia (Borneo).
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  • Like other langurs, Silvered Leaf Monkeys (Trachypithecus cristatus) are highly social and travel in cohesive groups numbering up to 40 or more individuals. They have a specialized diet, feeding almost exclusively on high-protein leaves. Sarawak, Malaysia (Borneo).
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  • This large male Northwest Bornean Orangutan (Pongo pygmaeus pygmaeus) is just beginning to develop cheek pads, a sign of maturity and placing his age to perhaps between 15 to 20 years. Batang Ai, Sarawak, Malaysia (Borneo).
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  • White-fronted Brown Lemur (Eulemur albifrons), male. Marojejy National Park, Antsiranana, Madagascar.
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  • The newly distinguished Kayan Loris (Nycticebus kayan) differs from other Bornean loris species by a combination of morphological traits. Sarawak, Malaysia (Borneo).
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  • A Northern Bamboo Lemur (Hapalemur occidentalis) in the foothills of Marojejy National Park, one of at least five species of Hapalemur distributed throughout the forests of Madagascar. True to their name, these small lemurs subsist on a diet that consists largely of young bamboo leaves, a remarkable feat considering the high cyanide content found in the plant’s tissues. It is still not completely understood exactly how these lemurs are able to survive on a diet so tainted with cyanide that it would kill animals much larger than themselves. Antsiranana, Madagascar.
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  • This large male Northwest Bornean Orangutan (Pongo pygmaeus pygmaeus) is just beginning to develop cheek pads, a sign of maturity and placing his age to perhaps between 15 to 20 years. Batang Ai, Sarawak, Malaysia (Borneo).
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  • Geoffroy’s Tamarin (Saguinus geoffroyi). Colón, Panama.
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  • The Sumatran Surili (Presbytis melalophos) is endemic to the island of Sumatra.  Like other leaf monkeys, this species is primarily folivorous, but also consumes flowers and fruits. Kerinci Seblat National Park, West Sumatra, Indonesia.
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  • Red-fronted Brown Lemur (Eulemur rufifrons). Ranomafana National Park, Madagascar.
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  • Threatened with encroaching cultivation and forest loss, a Southern Purple-Faced Langur (Trachypithecus vetulus vetulus) enters a tea plantation to feed on the fresh young leaves. These endangered monkeys are endemic to Sri Lanka’s southern wet forests, and although once widespread, are now facing severe habitat fragmentation due to roads and settlements. This can lead to human conflicts where the monkeys are inevitably forced to exploit agricultural areas for food. Sinharaja National Park. Sri Lanka.
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  • Silvered Leaf Monkey (Trachypithecus cristatus). Sarawak, Malaysia (Borneo).
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  • Southern Purple-faced Langur (Trachypithecus vetulus vetulus). Sinharaja National Park, Sri Lanka.
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  • Proboscis Monkey (Nasalis larvatus), male. Sarawak, Malaysia (Borneo).
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  • A large group of Proboscis Monkeys (Nasalis larvatus) prepares for sleep in a tree overhanging a river. Sabah, Malaysia (Borneo).
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  • The Maroon Langur (Presbytis rubicunda), also known as the Red Leaf Monkey, is endemic to Borneo and nearby Karimata Island, with several subspecies occurring throughout its range. Like other leaf monkeys they are almost completely arboreal and feed on foliage, seeds, and some fruits. Sabah, Malaysia.
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  • Young Ring-tailed Lemurs (Lemur catta) will transition from riding on the belly to the back of their mother after 4-6 weeks of age. Amboasary, Madagascar.
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  • Proboscis Monkey (Nasalis larvatus). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Black-and-white Ruffed Lemur (Varecia variegata), a Critically Endangered species and undoubtedly one of the most beautiful of all lemurs. Entirely dependent on tall undisturbed rainforests, these lemurs are threatened by habitat loss and are suffering from an island-wide population decline. Ranomafana National Park, Madagascar.
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  • A Maroon Langur (Presbytis rubicunda) pauses for a break while munching on his chosen leaf. Endemic to Borneo and a few offshore islands, these leaf monkeys feed on a wide range of foliage and unripe fruits, traveling in groups of up to a dozen individuals. One theory regarding their distinctive bright orange coloration is that this may help to confuse some predators into mistaking them for a much larger and stronger Orangutan, and thus deter attacks. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • The bright orange coloration of this infant Silvered Langur (Trachypithecus cristatus) helps them to be seen and hence guarded by the entire troop. They will gradually lose this coloration after 3 months of age. Bako National Park, Sarawak, Malaysia (Borneo).
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  • Verreaux's Sifaka (Propithecus verreauxi), mother with young. Amboasary, Madagascar.
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  • Northern Ceylonese Slender Loris (Loris lydekkerianus nordicus). North Western Province, Sri Lanka.
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  • The Maroon Langur (Presbytis rubicunda) is endemic to Borneo and nearby Karimata Island, with several subspecies occurring throughout its range. Like other leaf monkeys they are almost completely arboreal and feed on foliage, seeds, and some fruits. Sabah, Malaysia.
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  • Verreaux's Sifaka (Propithecus verreauxi). Amboasary, Madagascar.
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  • Rufous Mouse Lemur (Microcebus rufus), feeding on katydid. Ranomafana National Park, Madagascar.
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  • Brown Mouse Lemur (Microcebus rufus). Ranomafana National Park, Madagascar.
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  • Brown Mouse Lemur (Microcebus rufus). Ranomafana National Park, Madagascar.
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  • Verreaux's Sifaka (Propithecus verreauxi), crossing road. Popularly known as the ‘Dancing Sifakas’ for their remarkable hopping agility, few visitors realize that the dancing spectacle is in part due to the loss of these animals’ habitat: they naturally dwell in dry and spiny forests and only descend to the ground when they must cross a cleared area. Amboasary, Madagascar.
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  • Owston's Palm Civet (Chrotogale owstoni), a nocturnal carnivore from Laos and Vietnam, is a highly vulnerable species due to illegal poaching. Captive. Endangered Primate Rescue Centre, Cuc Phuong National Park, Vietnam.
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  • Owston's Palm Civet (Chrotogale owstoni), a nocturnal carnivore from Laos and Vietnam, is a highly vulnerable species due to illegal poaching. Captive. Endangered Primate Rescue Centre, Cuc Phuong National Park, Vietnam.
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  • Morning sun illuminates emergent baobab trees (Adansonia rubrostipa) in Madagascar's dry deciduous forest, one of the island's most endangered habitats. Despite receiving no rain for up to nine months each year, these forests are home to an astonishing diversity of wildlife, including Madam Berthe's Mouse Lemur, the world's smallest primate. Sadly, due to logging and land clearing, these forests have been reduced to less than 3% of their original extent, and now remain in only a few reserves along the western coast. Kirindy Reserve, Madagascar.
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  • By means of its extensive skin membranes, the Sunda Colugo (Galeopterus variegatus borneanus), is capable of gliding impressive distances between trees. This adult is carrying a young offspring on its underside. Bearing no relation to true lemurs, colugos are in their own unique order and are only distantly related to primates and treeshrews. They are nocturnal, feeding on leaves, flowers, and sap. Recent research on morphological and genetic variations indicate that the Bornean and Javan subspecies may merit recognition as distinct species. Sarawak, Malaysia (Borneo).
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  • With their vertical pupils and opposable thumbs, Phyllomedusa are easily some of the most distinguishable of all tree frogs. However, it is their peculiar form of locomotion that is perhaps their most striking feature: when seen walking slowly down a branch with their bodies raised, they appear decidedly ‘un-froglike’. Studies into their anatomy reveal that they possess a specialized musculature that not only allows them to move their limbs independently (as opposed to the simultaneous bilateral hopping motions of most other frogs), but also gives them a ‘precision grip’ which enables steady movement over thin branches. This grasping ability is rare among animals and is shared predominantly with higher primates, hence leading to one of their common names: ‘Monkey Frogs’. Coca, Ecuador.
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