Chien C. Lee

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  • The Chocoan Rainforest along the Pacific coast of Panama, Colombia, and northern Ecuador is one of the world's most endangered rainforest habitats. It is separated from the Amazon rainforest by the Andes mountains, and is home to a great diversity of endemic species. Esmeraldas, Ecuador.
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  • A wide swath of virgin rainforest cloaks the foothills of the Kobowre Mountains in western Papua. Indonesian New Guinea hosts some of the largest remaining tracts of primary tropical rainforest in the world.
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  • Virgin rainforest in western Borneo. West Kalimantan, Indonesia.
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  • Lowland rainforest in Masoala National Park. Antsiranana, Madagascar.
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  • Bornean rainforest trees generally have relatively shallow root systems because moisture is abundant and nutrients are most available near the surface of the soil. Reinforcement of tree's stability is provided in some species by conspicuous root buttresses. Sarawak, Malaysia (Borneo).
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  • Lowland rainforest in Masoala National Park. Antsiranana, Madagascar.
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  • Sunrise and mist over virgin rainforest in northern Borneo. Danum Valley Conservation Area, Sabah, Malaysia.
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  • Although the canopy of the Borneo rainforest is bathed in the heat of the equatorial sun, as little as 5 percent of sunlight reaches the forest floor. Sabah, Malaysia (Borneo).
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  • Morning sunlight scarcely penetrates the dense mist rising from virgin rainforest in the proposed Ulu Baleh National Park. This area is located deep in the remote interior of Sarawak, close to the geographic center of Borneo. Sarawk, Malaysia.
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  • A lone emergent Tapang (Koompassia excelsa) tree towers above the mist-shrouded canopy of the Borneo rainforest to catch the morning sun. This is one of the tallest tropical tree species with recorded heights of over 85 meters. Sabah, Malaysia.
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  • An aerial view reveals the thick mist rising from the canopy after heavy rains on virgin rainforest in northern Borneo. These forests often receive over 4000ml of rain annually. Sabah, Malaysia.
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  • This lowland rainforest in New Guinea is dominated by large Licuala fan palms. Papua, Indonesia.
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  • Large stretches of virgin rainforest still remain in the remote interior of eastern Borneo. East Kalimantan, Indonesia.
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  • Lowland rainforest in Esmeraldas, Ecuador.
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  • Lowland rainforest in Masoala National Park. Antsiranana, Madagascar.
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  • In the dense tropical rainforests of Borneo, plants struggle to compete for their share of the sunlight. As little as 2% of sunlight reaches the forest floor. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Once their camouflage has failed, many insects resort to an alternative strategy - trying to make themselves appear as large and intimidating as possible. This harmless leaf katydid (Tympanophyllum arcufolium) from the Borneo rainforest is attempting to do just that by standing on its head and spreading its translucent wings. Male Tympanophyllum katydids are renowned for the loud and eerie violin-like calls they make, which adds to the magical abience of the Bornean rainforest at night. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • View of canopy of lowland mixed dipterocarp forest during a mass flowering event. Sabah, Malaysia (Borneo).
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  • Hidden in the dense leaf litter of the rainforest floor, an Amazonian Horned Frog (Ceratophrys cornuta) lies in wait for its next meal. With a mouth wider than the length of its body, and a voracious appetite to boot, these frogs can consume prey as large as small reptiles and rodents. This species is widely distributed in the Amazon Basin, but nowhere particularly abundant. Yasuní National Park, Ecuador.
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  • A fan palm (Licuala valida) reaches towards the light in the understory of the Borneo rainforest. Sarawak, Malaysia (Borneo).
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  • The Long-tongued Nectar Bat (Macroglossus minimus) is an important pollinator of many rainforest trees. Here it is feeding on the nectar of banana flowers. Sarawak, Malaysia (Borneo).
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  • The Borneo rainforest understory is home to a great diversity of fan palms, with nearly 50 species recorded. Many, including this species (Licuala petiolulata), are endemic to the island. Sarawak, Malaysia (Borneo).
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  • A young Parson's Chameleon (Calumma parsonii) in the rainforest of Masoala National Park. Antsiranana, Madagascar.
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  • A giant leaf katydid (Pseudophyllus hercules), one of the world's largest, rests in the rainforest understory. Active only at night, they use their superb camouflage to remain undetected by predators during the day. Sarawak, Malaysia (Borneo).
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  • In one of the most magnificent remaining stands of virgin rainforest in northern Borneo, the Temburong River winds its way into the interior of the 500 sq. km. Ulu Temburong National Park in Brunei. With most of this tiny nation’s wealth coming from offshore oil, its forests have largely been spared from logging and oil palm plantations. Temburong, Brunei Darussalam.
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  • A delicate wild begonia (Begonia ambodiforahensis) from the ever-wet rainforest of northeastern Madagascar. Masoala National Park, Madagascar.
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  • A juvenile Oriental Vine Snake (Ahaetulla prasina) basks in a beam of sunlight in the rainforest of Tangkoko National Park. These highly arboreal snakes are excellent climbers and are adept at hunting lizards, their favored prey. North Sulawesi, Indonesia.
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  • A clear stream cascades down layered beds of sandstone amidst lowland rainforest. Samarakan, Sarawak, Malaysia (Borneo).
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  • Lowland rainforest on hilly terrain is cleared and terraced for planting of oil palm.
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  • Lowland rainforest on hilly terrain is cleared and terraced for planting of oil palm.
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  • The Long-tongued Nectar Bat (Macroglossus minimus) is an important pollinator of many rainforest trees including mangroves and bananas. Sarawak, Malaysia (Borneo).
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  • A large male Bornean Clouded Leopard (Neofelis diardi) is caught on camera trap as he patrols his territory in the rainforest of Danum Valley, Sabah. Borneo's largest cat species, the clouded leopard preys on a variety of smaller wildlife ranging from deer and wild boar to monkeys. Sabah, Malaysia (Borneo).
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  • With its varied topography of limestone mountains and lowland rainforest, Mulu National Park is home to one of the richest floral diversities in Borneo. Sarawak, Malaysia (Borneo).
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  • Spanning nearly 7 million square kilometers, it is difficult to conceptualize just how vast the Amazon rainforest is. Here, the Tiputini River winds its way through a tiny corner of the Amazon Basin in eastern Ecuador, eventually emptying its waters into Peru, Brazil, and then the Atlantic Ocean nearly 3 thousand kilometers away. Having lost nearly 20% of its forest cover in recent decades, the Basin is still under siege today from threats such as mining, oil drilling, and clearing for farming. Despite these losses, much still remains intact and just as imperative as ever to continue protective efforts for these invaluable forests. Yasuní National Park, Ecuador.
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  • A male Great Argus Pheasant (Argusianus argus) is one of the most magnificent birds in the Bornean rainforest. Despite its large size it is shy and seldom seen, though its haunting call is frequently heard. Sabah, Malaysia (Borneo).
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  • Fruit and nectar bats (family Pteropodidae) play an important role in the dispersal and pollination of many rainforest trees including figs, bananas, and durian. Over 17 species occur in Borneo, including the widespread Short-nosed Fruit Bat (Cynopterus brachyotis, pictured here). Unlike insectivorous bats, most Pteropodids do not echolocate and instead rely on their large eyes and keen sense of smell for nocturnal navigation. Sarawak, Malaysia (Borneo).
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  • Seeds of Alsomitra macrocarpa, a large tropical climbing vine, lie scattered on the forest floor.  Each seeds is adorned with a large transparent wing which enables it to glide slowly down from the rainforest canopy, sometimes traveling hundreds of yards. Sarawak, Malaysia (Borneo).
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  • A tiny bush frog (Philautus cf. aurantium) calls from its perch in the Bornean rainforest. Sarawak, Malaysia (Borneo).
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  • After quickly selecting a ripe fig (Ficus fistulosa), a Short-nosed Fruit Bat (Cynopterus brachyotis) makes off with her prize to dine some distance away at a safe perch, thereby helping to disperse the tree's seeds that will ultimately lead to fruit for her future generations. Figs reach their pinnacle of diversity in Borneo with at least 150 species and, perhaps more than any other group of plants, are considered keystone species of the rainforest because of the complex interdependences they exhibit with countless animals and insects. Sarawak, Malaysia (Borneo).
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  • A giant bloom of Rafflesia arnoldii, the largest flower in the world, unfolds its petals in the Sumatran rainforest.  It takes nealy 2 days for the flower to fully open. West Sumatra, Indonesia.
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  • A juvenile leaf insect (Nanophyllium australianum) from the rainforest of Iron Range National Park. Queensland, Australia.
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  • An unusually-shaped epiphytic orchid (Bulbophyllum uniflorum) blooms on a mossy tree in Sumatra's montane rainforest. West Sumatra, Indonesia.
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  • Madagascar Fire Millipede (Aphistogoniulus hova), found in the rainforest of Masoala National Park. Antsiranana, Madagascar.
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  • The luminescent nature of a cluster of seemingly ordinary mushrooms (Favolaschia manipularis) becomes revealed in the darkening dusk of the Borneo rainforest. Sabah, Malaysia (Borneo).
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  • A shy bird of the rainforest floor, the beautiful Blue-headed Pitta (Hydrornis baudii) is endemic to Borneo. Danum Valley Conservation Area, Sabah, Malaysia.
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  • A Lined Gecko (Gekko vittatus) perches in its rainforest habitat on the island of Halmahera, central Indonesia. Sometimes referred to as the "Skunk Gecko" because of its distinctive coloration, these are popular animals in the pet trade and large numbers are collected and exported yearly from Indonesia and the Solomon Islands. North Maluku, Indonesia.
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  • Established in 1991, Ranomafana National Park protects over 160 sq. mi. of tropical rainforest, a habitat which has been reduced to less than 10% of its original extent in Madagascar.
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  • Seeds of Alsomitra macrocarpa, a large tropical climbing vine, lie scattered on the forest floor.  Each seeds is adorned with a large transparent wing which enables it to glide slowly down from the rainforest canopy, sometimes traveling hundreds of yards. Sarawak, Malaysia (Borneo).
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  • The Malay Civet (Viverra tangalunga) is a shy nocturnal animal of the rainforest.  This individual was photographed by means of an infrared camera trigger. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • A stream through lowland rainforest. Mulu National Park, Sarawak (Borneo).
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  • Hidden in the dense leaf litter of the rainforest floor, an Amazonian Horned Frog (Ceratophrys cornuta) lies in wait for its next meal. With a mouth wider than the length of its body, and a voracious appetite to boot, these frogs can consume prey as large as small reptiles and rodents. This species is widely distributed in the Amazon Basin, but nowhere particularly abundant. Yasuní National Park, Ecuador.
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  • Hidden in the dense leaf litter of the rainforest floor, an Amazonian Horned Frog (Ceratophrys cornuta) lies in wait for its next meal. With a mouth wider than the length of its body, and a voracious appetite to boot, these frogs can consume prey as large as small reptiles and rodents. This species is widely distributed in the Amazon Basin, but nowhere particularly abundant. Yasuní National Park, Ecuador.
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  • Goniothalamus ridleyi. Cauliflory (flowering and fruiting along the trunk) is a common phenomenon among many rainforest tree species, and is an adaptation to reach pollinating and dispersing animals in the understory. East Kalimantan, Indonesia (Borneo).
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  • A juvenile leaf insect (Nanophyllium australianum) from the rainforest of Iron Range National Park. Queensland, Australia.
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  • Polyalthia cauliflora. Cauliflory (flowering and fruiting along the trunk) is a common phenomenon among many rainforest tree species and is an adaptation to reach pollinating and dispersing animals in the understory. Sarawak, Malaysia (Borneo).
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  • The Bornean Forest Dragon (Gonocephalus bornensis) is a large arboreal agamid lizard endemic to the rainforests of Borneo. It feeds on ants and spiders. Sarawak, Malaysia (Borneo).
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  • Named in honor of singer-songwriter David Bowie, this large huntsman spider (Heteropoda davidbowie) occurs in rainforests from Thailand to Sumatra and Borneo.
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  • The enormous Palm Cockatoo (Probosciger aterrimus), sometimes called the Goliath Cockatoo, is one of the largest parrots in Asia. It is found in the rainforests of New Guinea and northern Australia. West Papua, Indonesia (New Guinea).
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  • The Palawan Peacock Pheasant (Polyplectron napoleonis) is a shy ground-dwelling bird endemic to the rainforests the island of Palawan, Philippines.
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  • Found throughout the lowland rainforests of Sarawak, the loud honking call of the Bornean Horned Frog (Pelobatrachus nasutus) is commonly heard before the onset of heavy thunderstorms. When disturbed it remains completely motionless, relying on its excellent camouflage to escape detection from potential predators. Sabah, Malaysia (Borneo).
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  • The Spotted Cuscus (Spilocuscus maculatus chrysorrhous) is a widespread arboreal marsupial occurring in the lowland rainforests of New Guinea and northern Australia. It is solitary and nocturnal, feeding primarily on leaves and fruits. Central Papua, Indonesia (New Guinea).
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  • The Brown Spiny Rat (Maxomys rajah) is found in lowland rainforests where it forages for food on the ground and in low vegetation. Sarawak, Malaysia (Borneo).
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  • An increasingly rare sight, clear rocky streams are an important and indicative component of pristine habitats in Borneo rainforests. Sarawak, Malaysia.
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  • The Spotted Cuscus (Spilocuscus maculatus chrysorrhous) is a widespread arboreal marsupial occurring in the lowland rainforests of New Guinea and northern Australia. It is solitary and nocturnal, feeding primarily on leaves and fruits. Central Papua, Indonesia (New Guinea).
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  • Found throughout the lowland rainforests of Sarawak, the loud honking call of the Bornean Horned Frog (Pelobatrachus nasutus) is commonly heard before the onset of heavy thunderstorms.  When disturbed it remains completely motionless, relying on its excellent camouflage to escape detection from potential predators. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Found throughout the lowland rainforests of Sarawak, the loud honking call of the Bornean Horned Frog (Pelobatrachus nasutus) is commonly heard before the onset of heavy thunderstorms.  When disturbed it remains completely motionless, relying on its excellent camouflage to escape detection from potential predators. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Red-crowned Barbet (Psilopogon rafflesii), a frugivorous bird from the rainforests of Southeast Asia. Sarawak, Malaysia (Borneo).
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  • The remarkable feathered antennae of a Callirhipid cedar beetle (Callirhipis sp.) from the montane rainforests of New Guinea. Papua, Indonesia.
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  • A male harvestman (Quindina limbata) within a nest he has constructed on the side of a fallen log. He will remain in this nest for several months to mate with visiting females and guard the eggs which have been buried within the mud floor. Limón, Costa Rica.
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  • The Rufous Piculet (Sasia abnormis) is the smallest woodpecker in Borneo.  Here a bird is making its territorial call by drumming on a spiny rattan petiole. Sabah, Malaysia (Borneo).
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  • A stream cascades over sandstone layers in Lambir Hills National Park. Sarawak, Malaysia (Borneo).
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  • A male Bornean Tree-hole Frog (Metaphrynella sundana) calls for a mate from its lair.  These frogs are able to modify the pitch of their call to match the resonating frequency of the tree-hole chamber, thereby amplifying their volume. Sarawak, Malaysia (Borneo).
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  • Leaf katydid (Sathrophylliopsis longepilosa), female. Sarawak, Malaysia (Borneo).
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  • Variable Sticky Frog (Kalophrynus heterochirus). Like other species in the genus, this frog secretes a sticky substance when disturbed. Sarawak, Malaysia (Borneo).
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  • Keel-bellied Vine Snake (Dryophiops rubescens), a lizard-eating arboreal colubrid. Sarawak, Malaysia (Borneo).
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  • Tiger Leech (Haemadipsa sp. aff. picta). Sabah, Malaysia (Borneo).
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  • Sabah Bamboo Pitviper (Trimeresurus sabahi). Sarawak, Malaysia (Borneo).
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  • Flat-nosed Pitviper (Craspedocephalus puniceus). West Java, Indonesia.
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  • The Black-bearded Tomb Bat (Taphozous melanopogon) roosts in caves and large rock crevices, emerging at night to feed on insects high above the forest canopy. It is found throughout much of mainland Southeast Asia as well as Borneo, Sumatra, and Java. It's echolocation calls while flying are audible as high pitched clicks. Sarawak, Malaysia (Borneo).
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  • A crystal-clear stream flows through the forest at Khao Nor Chuchi Wildlife Sanctuary.  These spring-fed waters which emerge from limestone rock are highly alkaline and contain high amounts of dissolved calcium carbonate. Krabi, Thailand.
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  • A male Whitehead's Broadbill (Calyptomena whiteheadi) returns to its nest where it passes food to the hungry chick. Both male and female care for the nestlings, providing insects and fruit, as well as removing fecal pellets. This scarcely-seen species is the largest of all the green broadbills (reaching 27cm in length) and is endemic to the montane forests of Borneo. Sabah, Malaysia.
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  • Threatened by habitat loss and hunting pressure, the Crested Black Macaque (Macaca nigra nigra) is now listed as Critically Endangered. This species lives only in forested areas at the northernmost tip of the island of Sulawesi. North Sulawesi, Indonesia.
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  • Endemic to the mountains of central Sulawesi, the stream-breeding Loka Flying Frog (Rhacophorus monticola) is highly variable in coloration. South Sulawesi, Indonesia.
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  • This Collared Nightjar (Gactornis enarratus), a Madagascar endemic, hides itself among the leaf litter of the forest floor while it sleeps. Placed in its own genus, this species is distinct from all other nightjars (family Caprimulgidae) by its plumage, eggs, and behavior. Mantadia National Park, Madagascar.
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  • Wild nutmeg (Myristica sp.), fruits. Sarawak, Malaysia (Borneo).
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  • Extremely agile on the wing, the Bat Hawk (Macheiramphus alcinus) is a specialist predator of bats and swiftlets, often waiting near cave entrances for its prey to emerge. Sarawak, Malaysia (Borneo).
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  • Bronzeback (Dendrelaphis terrificus). Gorontalo, Sulawesi, Indonesia.
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  • An enormous pitcher trap (Nepenthes rafflesiana) rests on the forest floor, an open but sinister invitation for wandering insects. This carnivorous plant has been known to occasionally catch and consume mice in its voluminous pitchers which can reach over 40cm in height. Belait, Brunei Darussalam (Borneo).
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  • Batang Ai National Park. Sarawak, Malaysia (Borneo).
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  • Hikida's Forest Dragon (Hypsilurus hikidanus). Central Papua, Indonesia (New Guinea).
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  • Trilobite beetle (Platerodrilus foliaceus), female. East Kalimantan, Indonesia (Borneo).
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  • Although the Banded Coral Snake (Calliophis intestinalis) is not aggressive, it possesses an extremely toxic venom like other Elapids. When disturbed it displays its red tail and the brightly colored bands on its undersurface which serve as a warning signal to would-be predators. Sarawak, Malaysia (Borneo).
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  • Two Giant Forest Ants (Dinomyrmex gigas) from different colonies confront each other in a territorial dispute.  Rather than engage in full combat, this species usually resolves such conflict in a series of ritualistc fights. Sarawak, Malaysia (Borneo).
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  • Paradise Tree Snake (Chrysopelea paradisi). Sarawak, Malaysia (Borneo).
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  • Lantern Bug (Pyrops candelarius) from Indochina.  Like other planthoppers, this insect feeds on tree sap by means of its piercing mouthparts.  The name refers to the erroneous belief that the snout-like projection emits light. Cuc Phuong National Park, Vietnam.
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  • Geometrid moth (Agathia deliciosa). Sarawak, Malaysia (Borneo).
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  • This small dung beetle (Paragymnopleurus maurus) is preparing a ball made from monkey droppings in which it will deposit its eggs and then bury. Sarawak, Malaysia (Borneo).
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  • Bagworm caterpillars (moths of the family Psychidae) are known for their habit of hiding themselves in a portable shelter of plant materials that they secure with silken threads, with each species having its own preference of design. This species begins with small accurately trimmed twigs formed in a 3-sided spiral pyramid, and adds additional larger twigs at the end as the caterpillar grows in size, in a masterful example of miniature craftsmanship. East Kalimantan, Indonesia (Borneo).
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  • Undoubtedly one of the world's most remarkable plants, the parasitic Rafflesia holds the record for having the largest flower, sometimes reching over one meter across. Their macabre beauty belies a clever deception: the dark reddish coloration and putrid stench of the bloom mimic the rotting carcass of an animal, which thus lures carrion flies into the interior chamber where they will pollinate it. There are around 30 species in the genus, with variable flower sizes, this being R. tuan-mudae from Borneo, one of the largest. Sarawak, Malaysia (Borneo).
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  • Crab spiders of the genus Amyciaea are excellent mimics of Weaver Ants (Oecophylla), their preferred prey. In this manner they are able to both avoid predators and easily approach within striking distance of the ants. North Maluku, Indonesia.
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