Chien C. Lee

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  • Dung beetle (Paragymnopleurus sparsus), rolling scat of Sambar Deer. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Dung Beetle (Epilissus splendidus). Mantadia National Park, Madagascar.
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  • Like other members of its genus, the males of this rhinoceros beetle (Xylotrupes ulysses clinias) exhibit variation in horn size with some individuals having very long horns (as shown here) and others with very short horns. This extreme dimorphism is controlled by an environmentally influenced genetic switch. South Sulawesi, Indonesia.
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  • A large male rhinoceros beetle (Beckius beccarii). This species is endemic to the forested regions of New Guinea. West Papua, Indonesia.
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  • Dung beetle (Proagoderus watanabei), male feeding on herbivore scat. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Dung beetle (Scarabaeus sp.), rolling dung. Bundala National Park, Sri Lanka.
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  • This small dung beetle (Paragymnopleurus maurus) is preparing a ball made from monkey droppings in which it will deposit its eggs and then bury. Sarawak, Malaysia (Borneo).
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  • Dung beetle (Paragymnopleurus maurus), rolling scat of Sambar Deer. Danum Valley, Sabah, Malaysia (Borneo).
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  • Looking as if it were dipped in liquid gold, a jewel scarab (Chrysina argenteola) almost seems more mineral than insect. Renowned for their remarkable colorations, Chrysina beetles fetch high prices among collectors, with some species and color variations fetching hundreds of dollars. Recent research into the metallic nature of their exoskeletons reveals that it possesses unique properties in the way it reflects polarized light, essentially being 'optically ambidextrous'. The reason why they have such unusual colorations still remains a mystery, although it has been suggested that the highly reflective surface may serve to camouflage them under the right lighting conditions, or even perhaps dazzle predators. Canandé Reserve, Esmeraldas, Ecuador.
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  • A male flower scarab (Mecynorhina harrisi), prepared to ward off rivals with his formidable horn, guards a female while she feeds on tree sap. Udzungwa Mountains, Tanzania.
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  • A pair of the largest dung beetles in Borneo (Catharsius dayacus) clean up after the droppings of a Sambar Deer in the rainforest of Danum Valley, Sabah. Although this species does not roll balls like its more well-known relatives, it nevertheless is responsible for burying large amounts of animal scat and provides a vital ecological service. Nearly a hundred species of dung beetles have been recorded from Danum alone; studies have shown that their diversity not only directly reflects the wide assortment of vertebrates in an area but also can serve as a valuable indicator of habitat heterogeneity and forest disturbance. Sabah, Malaysia (Borneo).
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  • One of the largest beetles in Borneo, this magnificent species (Chalcosoma moellenkampi) is found in lowland and hilly rainforests and is frequently attracted to lights. Only the males have the distinctive horns. Danum Valley, Sabah, Malaysia (Borneo).
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  • Marauder Ants (Carebara affinis) attacking a beetle grub. The minor workers are assisted by their larger sisters, the majors and super-majors, which use their powerful mandibles to cut the grub into smaller pieces. Sarawak, Malaysia (Borneo).
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  • Marauder Ants (Carebara affinis) attacking a beetle grub. The minor workers are assisted by their larger sisters, the majors and super-majors, which use their powerful mandibles to cut the grub into smaller pieces. Sarawak, Malaysia (Borneo).
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