Chien C. Lee

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  • Endemic to the mountains of central Sulawesi, the stream-breeding Loka Flying Frog (Rhacophorus monticola) is highly variable in coloration. South Sulawesi, Indonesia.
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  • Larut Bush Frog (Philautus larutensis), male. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Penan Flying Frog (Leptomantis penanorum), a very rare species known from only a few specimens. This is the first photograph of a female ever recorded for this species. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Ryabov's Bug-eyed Frog (Theloderma ryabovi), female. Thua Thien Hue, Vietnam.
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  • Peacock Gliding Frog (Rhacophorus kio). Yunnan, China.
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  • Wallace's Flying Frog (Rhacophorus nigropalmatus) is one of the largest of all tree frogs in Borneo. It is capable of gliding down from the forest canopy by using its enlarged webbed feet as parachutes. Sabah, Malaysia (Borneo).
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  • Bornean Opposite-fingered Tree Frog (Feihyla inexpectata), first photographic record for this species in Sarawak. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • The Warted Frog is a rare species in Borneo, known from only a handful of sightings in the northwest. It is presumably a tree-hole breeder as in other Theloderma. Sarawak, Malaysia (Borneo).
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  • Belalong Tree Frog (Leptomantis belalongensis), male. Only recently described, this tiny arboreal frog is endemic to just a few river basins in northwestern Borneo. Ulu Temburong National Park, Brunei (Borneo).
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  • Jade Tree Frog (Zhangixalus dulitensis). Ulu Temburong National Park, Brunei (Borneo).
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  • South-Vietnamese Bug-Eyed Frog (Theloderma vietnamense). Thua Thien Hue, Vietnam.
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  • Dring's Bush Frog (Philautus juliandringi), male. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Perhaps one of the most beautiful of all Borneo's frogs: the Borneo Flying Frog (Rhacophorus borneensis). These gliding amphibians spend most of their life in the tree canopy, only rarely descending to ground level. Sarawak, Malaysia (Borneo).
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  • Perhaps one of the most beautiful of all Borneo's frogs: the Borneo Flying Frog (Rhacophorus borneensis). These gliding amphibians spend most of their life in the tree canopy, only rarely descending to ground level. Sarawak, Malaysia (Borneo).
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  • Near the summit of Gunung Murud (Sarawak's highest mountain), an newly described species of tiny bush frog (Philautus nepenthophilus) hides within the fluid of a carnivorous pitcher plant (Nepenthes mollis), apparently unaffected by the plant's digestive juices therein. Phytotelmata (water bodies held by plants) provide living quarters and breeding grounds for many unique creatures which are completely dependent on them. Pulong Tau National Park, Sarawak, Malaysia (Borneo).
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  • Distinguished by its green/yellow eyes, Hose's Bush Frog (Philautus hosii) is endemic to Borneo and considered Near Threatened due to severe habitat loss. Adults are usually found perched on vegetation over streams in lowland rainforest. Sarawak, Malaysia (Borneo).
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  • File-eared Tree Frog (Polypedates otilophus). One of the largest tree frogs in Borneo, this species is named for the sharp ridge behind its eye, the function of which is unknown. It breeds in stagnant pools in lowland rainforest. Sarawak, Malaysia (Borneo).
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  • A Harlequin Gliding Frog (Rhacophorus pardalis) struggles futilely as it is being slowly swallowed alive by a Black-Headed Cat Snake (Boiga nigriceps). Stagnant pools of water in the Borneo rainforest serve as important breeding grounds for many tree frogs which gather around in great numbers, a fact that some predators wisely take advantage of. Kubah National Park, Sarawak, Malaysia (Borneo).
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  • Wallace’s Flying Frog (Rhacophorus nigropalmatus) is one of the largest of all tree frogs in Borneo. It is capable of gliding down from the forest canopy by using its enlarged webbed feet as parachutes. Sarawak, Malaysia (Borneo).
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  • Bornean Gliding Frog (Rhacophorus borneensis), female. Sarawak, Malaysia (Borneo).
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  • The enlarged webbed feet of the Harlequin Tree Frog (Rhacophorus pardalis) enable it to maneuver in mid-air and slow its descent from a higher perch towards a safe landing. Sarawak, Malaysia (Borneo).
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  • A bush frog (Philautus amoenus) emerges from it's watery shelter in the pitcher of Nepenthes × harryana. Mount Kinabalu. Sabah, Malaysia (Borneo).
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  • Wallace's Flying Frog (Rhacophorus nigropalmatus), male. Sarawak, Malaysia (Borneo).
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  • Wallace's Flying Frog (Rhacophorus nigropalmatus), male. Sarawak, Malaysia (Borneo).
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  • Ryabov's Bug-eyed Frog (Theloderma ryabovi), female. Thua Thien Hue, Vietnam.
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  • Bornean Gliding Frog (Rhacophorus borneensis), pair in amplexus. Danum Valley, Sabah, Malaysia (Borneo).
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  • Red-legged Frog (Leptomantis rufipes). Sarawak, Malaysia (Borneo).
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  • Harlequin Gliding Frog (Rhacophorus pardalis). Sarawak, Malaysia (Borneo).
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  • Bush frog (Philautus nepenthophilus) in pitcher plant (Nepenthes mollis). Pulong Tau National Park, Sarawak, Malaysia (Borneo).
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  • File-eared Tree Frog (Polypedates otilophus). Danum Valley, Sabah, Malaysia (Borneo).
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  • Long-legged Bush Frog (Philautus kakipanjang). Sarawak, Malaysia (Borneo).
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  • Cinnamon Tree Frog (Nyctixalus pictus). Sarawak, Malaysia (Borneo).
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  • The dainty White-eared Tree Frog (Feihyla kajau) is endemic to the rainforests of Borneo. Sarawak, Malaysia.
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  • Unlike the rest of the Philippines, Palawan has a close biogeographic affinity with Borneo, which is separated by a shallow sea channel.  This is exemplified by many closely-related species of plants and animals, including Everett's Tree Frog (Philautus everetti), an inhabitant of hill and montane forests.
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  • Wallace's Flying Frog (Rhacophorus nigropalmatus). Sarawak, Malaysia (Borneo).
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  • Bornean Gliding Frog (Rhacophorus borneensis), female. Sarawak, Malaysia (Borneo).
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  • Kerangas Bush Frog (Philautus kerangae). Sarawak, Malaysia (Borneo).
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  • Bush frog (Philautus nepenthophilus). Pulong Tau National Park, Sarawak, Malaysia (Borneo).
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  • Golden-legged Bush Frog (Philautus aurantium), male. Sabah, Malaysia (Borneo).
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  • The White-eared Tree Frog (Feihyla kajau) lays its eggs in clusters on leaves overhanging small streams. When the tadpoles emerge they drop down into the water below. Sarawak, Malaysia (Borneo).
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  • File-eared Tree Frog (Polypedates otilophus). Sarawak, Malaysia (Borneo).
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  • Dark-eared Tree Frog (Polypedates macrotis). Sarawak, Malaysia (Borneo).
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  • A tiny bush frog (Philautus cf. aurantium) calls from its perch in the Bornean rainforest. Sarawak, Malaysia (Borneo).
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  • Tree frog (Leptomantis cf. penanorum). Sarawak, Malaysia (Borneo).
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  • Harlequin Tree Frog (Rhacophorus pardalis). Sabah, Malaysia (Borneo).
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  • Harlequin Tree Frog (Rhacophorus pardalis).  A female making a foam nest on a small tree overhanging a breeding pond.  After depsiting her eggs they will be fertilized by the two accompanying smaller males. Sarawak, Malaysia (Borneo).
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  • The Gading Flying Frog (Leptomantis gadingensis), described as recently as 2005, is a small species known from only a few localities in western Borneo.  It breeds in lowland streams. Sarawak, Malaysia (Borneo).
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  • Mossy Tree Frog (Philautus macroscelis), male. Sarawak, Malaysia (Borneo).
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  • Bush frog (Pseudophilautus sp.). Sinharaja National Park, Sri Lanka.
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  • Bush frog (Philautus nepenthophilus) in pitcher plant (Nepenthes mollis). Pulong Tau National Park, Sarawak, Malaysia (Borneo).
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  • Lanjak Bush Frog (Philautus refugii). Sarawak, Malaysia (Borneo).
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  • The mountains of Vietnam are home to several species of amphibians which are perfectly adapted for camouflage in the mossy cloud forests. Perched atop a small leaf, this Tiny Bubblenest Frog (Gracixalus supercornutus) can easily be mistaken for a clump of moss epiphylls. Bach Ma National Park, Vietnam.
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  • Borneo Opposite-fingered Tree Frog (Feihyla inexpectata), male. Sabah, Malaysia (Borneo).
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  • Bush frog (Philautus sp.). Sarawak, Malaysia (Borneo).
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  • The tiny Obscure Bush Frog (Philautus tectus) is endemic to Borneo where it can be found near small streams and seeps in lowland rainforest. This species has been listed as Vulnerable because it is limited to small scattered populations, many of which have been lost due to land development. Sarawak, Malaysia (Borneo).
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  • White-eared Tree Frog (Feihyla kajau), male. Sarawak, Malaysia (Borneo).
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  • Dull-green Shrub Frog (Pseudophilautus viridis). Central Province, Sri Lanka.
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  • A pair of Bornean Gliding Frogs (Rhacophorus borneensis) in amplexus. Like other Rhacophorid tree frogs, the female will deposit a mass of eggs on a leaf overhanging a pool of water which are then fertilized by the male. Sarawak, Malaysia (Borneo).
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  • A pair of Short-nosed Tree Frogs (Leptomantis gauni) in amplexus, perched on the foliage of a tree on the bank of a clear stream. Eggs will be deposited in a foam nest overhanging the water so that the emerging tadpoles can drop directly into the stream below. Sarawak, Malaysia (Borneo).
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