Chien C. Lee

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  • Without seeing it move, you could almost mistake the Golden Mantella (Mantella aurantiaca) for a bright orange plastic toy. This is one of Madagascar’s most endangered amphibians and is an icon for conservation of the island’s threatened wildlife. Efforts targeted at protecting this frog’s habitat, coupled with ex-situ breeding programs and reintroduction have helped to protect it from extinction in the wild, but it remains highly threatened and is still known to exist at only two small isolated patches of rainforest. Andasibe, Madagascar.
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  • Without seeing it move, you could almost mistake the Golden Mantella (Mantella aurantiaca) for a bright orange plastic toy. This is one of Madagascar’s most endangered amphibians and is an icon for conservation of the island’s threatened wildlife. Efforts targeted at protecting this frog’s habitat, coupled with ex-situ breeding programs and reintroduction have helped to protect it from extinction in the wild, but it remains highly threatened and is still known to exist at only two small isolated patches of rainforest. Andasibe, Madagascar.
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  • Threatened by habitat loss and hunting pressure, the Crested Black Macaque (Macaca nigra nigra) is now listed as Critically Endangered. This species lives only in forested areas at the northernmost tip of the island of Sulawesi. North Sulawesi, Indonesia.
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  • Only distantly related to other mongooses, the Narrow-striped Mongoose (Mungotictis decemlineata) is actually a member of the family Eupleridae and descendant from the same lineage as the much larger Fossa. Active by day, these small predators are endemic to the dry deciduous forests of western Madagascar where they live in small matriarchal family groups. Having only a patchy distribution, they are severely threatened by habitat loss. Kirindy Reserve, Madagascar.
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  • A juvenile Parson's Chameleon (Calumma parsoni). More frequently occurring in shades of green, this is one of the world's largest chameleons, with adult specimens sometimes reaching the size of a house cat. They are endemic to rainforest habitats on Madagascar and are threatened by both habitat loss and collection for the international pet trade. Andasibe, Madagascar.
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  • With its rough skin texture and mottled green coloration, it’s easy to see how the Mossy Rain Frog (Pristimantis museosus) is well equipped to blend in with the damp forests in which it dwells. This frog is endemic to the mountains of central Panama, a region which has seen drastic declines in many amphibians over the past few decades due to the introduction of the virulent chytrid fungus. Like other species of Pristimantis, the females of P. museosus lay large eggs that do not require water – the tadpoles transform into froglets entirely within the egg itself. This adaptation may have provided this species with some protection against chytrid, since the fungus most easily transfers among those species which breed in bodies of water. Nevertheless, this frog remains highly threatened and has disappeared from multiple sites across its range. Panamá Oeste, Panama.
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  • Threatened with encroaching cultivation and forest loss, a Southern Purple-Faced Langur (Trachypithecus vetulus vetulus) enters a tea plantation to feed on the fresh young leaves. These endangered monkeys are endemic to Sri Lanka’s southern wet forests, and although once widespread, are now facing severe habitat fragmentation due to roads and settlements. This can lead to human conflicts where the monkeys are inevitably forced to exploit agricultural areas for food. Sinharaja National Park. Sri Lanka.
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  • Endemic to the Bird's Head Peninsula of western New Guinea, the Vogelkop Tree Kangaroo (Dendrolagus ursinus) has become increasingly threatened by habitat loss and hunting. West Papua, Indonesia (New Guinea).
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  • A jewel of Ecuador’s high-elevation cloud forests is the spectacular Plate-billed Mountain Toucan (Andigena laminirostris). Unlike the more familiar toucans of lowland rainforests, these birds inhabit the coldest and wettest mountains of the Andes. Despite their predilection for these remote habitats, they are threatened by both habitat loss and capture for the pet trade. Mindo, Ecuador.
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  • One of the world's most endangered primates, the Bornean Banded Langur (Presbytis chrysomelas chrysomelas) has been threatened by both habitat loss and hunting. Now, only a few isolated populations remain in coastal Sarawak, with an estimated total of just a few hundred individuals. They feed on foliage and fruits in lowland rainforest. Sarawak, Malaysia (Borneo).
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  • Black-and-white Ruffed Lemur (Varecia variegata), a Critically Endangered species and undoubtedly one of the most beautiful of all lemurs. Entirely dependent on tall undisturbed rainforests, these lemurs are threatened by habitat loss and are suffering from an island-wide population decline. Ranomafana National Park, Madagascar.
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  • Endemic to the Bird's Head Peninsula of western New Guinea, the Vogelkop Tree Kangaroo (Dendrolagus ursinus) has become increasingly threatened by habitat loss and hunting. West Papua, Indonesia (New Guinea).
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  • Spectacled Flying Fox (Pteropus conspicillatus). Waigeo Island, West Papua, Indonesia.
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  • Sri Lankan Grey Langur (Semnopithecus priam thersites). Yala National Park, Sri Lanka.
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  • Nycticebus menagensis. One of four species of Slow Loris found in Borneo. All Slow Loris are nocturnal arboreal primates which feed on small animals, insects and soft fruits. Sarawak, Malaysia (Borneo).
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  • Endemic to northern Borneo, the Black-and-crimson Pitta (Pitta ussheri) is a shy forest bird which feeds on worms and other terrestrial invertebrates. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Endemic to the mountains of central Sulawesi, the stream-breeding Loka Flying Frog (Rhacophorus monticola) is highly variable in coloration. South Sulawesi, Indonesia.
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  • Kerangas Bush Frog (Philautus kerangae). Sarawak, Malaysia (Borneo).
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  • Pitcher plant (Nepenthes sibuyanensis). This large pitcher plant is endemic to the ultramafic Mount Guiting-guiting on Sibuyan Island, Philippines.
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  • With fewer than a thousand individuals remaining in the wild, the Critically Endangered Golden Bamboo Lemur (Hapalemur aureus) holds on to a tenuous existence in the rainforests of Madagascar. It’s discovery in the mid-1980’s was a crucial factor that led to the foundation of the now famous Ranomafana National Park – a reserve that protects over 400 sq. km. of tropical forest. Like other bamboo lemurs, it feeds primarily on bamboo shoots which, due to their high cyanide content, are lethally toxic to most other animals. Ranomafana National Park, Madagascar.
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  • Madagascar Plover (Charadrius thoracicus). Toliara, Madagascar.
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  • Marañón White-fronted Capuchin (Cebus yuracus). Yasuní National Park, Ecuador.
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  • Sulawesi Babirusa (Babyrousa celebensis). Gorontalo, Sulawesi.
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  • One of Borneo's most famous endemic mammals, the Proboscis Monkey (Nasalis larvatus) is restricted to coastal swamp forests and mangroves. Bako National Park, Sarawak, Malaysia (Borneo).
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  • Once considered one of the three most elusive birds in the world, the Long-tailed Ground Roller (Uratelornis chimaera) is a prized sighting for any birdwatcher. The entire family of Ground Rollers (Brachypteraciidae, 5 species) is endemic to Madagascr, but while most are inhabitants of rainforest, this is the only species found in in the arid spiny desert of the island’s extreme southwest. Even within this region it has as very restricted distribution and occurs at low densities, with only a few birds per square kilometer. Adapted for running swiftly on the ground with their powerful legs, these birds hunt for insects and small animals in the sand and leaf-litter. Toliara, Madagascar.
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  • Blue-legged Mantella (Mantella expectata), found only from a few locations in Madagascar's arid southwest. Isalo National Park, Madagascar.
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  • One of the most beautiful of all pitcher plants, Nepenthes edwardsiana produces enormous cylindrical pitchers which are strikingly colored. It is endemic to the Mount Kinabalu region in northern Borneo.
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  • Mountain Treeshrew (Tupaia montana) feeding at Nepenthes rajah.  Recent research has shown the the world's largest pitcher plant Nepenthes rajah is not exclusively carnivorous. Like the related N. lowii, this species attracts treeshrews by secreting nectar on the undersurface of the lid. These animals frequently leave their droppings in the pitcher, which serves as a valuable nitrogen source in their impoverished mountain habitat. Mount Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • An adult male Silvered Leaf Monkey (Trachypithecus cristatus) keeps a watchful eye out for any threats to his harem of females. Sarawak, Malaysia (Borneo).
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  • Leopard (Panthera pardus), juvenile. Ngamiland, Botswana.
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  • The least known of all of Sulawesi's monkeys, Heck's Macaque (Macaca hecki) occurs on the northwestern portion of the island. Five other endemic species of macaque are distributed in different regions of Sulawesi, occasionally producing hybrids where their ranges overlap. All are diurnal and feed largely on fruits. Central Sulawesi, Indonesia.
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  • The shy Maleo (Macrocephalon maleo) is a large megapode bird endemic to Sulawesi. They do not build nests but instead bury their eggs in huge pits dug in hot volcanic sand. The precocious young are able to fly and fend for themselves immediately after hatching. Central Sulawesi, Indonesia.
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  • Sharp-nosed Chameleon (Kinyongia oxyrhina), male. Udzungwa Mountains, Tanzania.
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  • Usambara Eyelash Viper (Atheris ceratophora). Udzungwa Mountains, Tanzania.
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  • Coronated Tree Frog (Triprion spinosus). Cartago, Costa Rica.
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  • Bornean Orangutan (Pongo pygmaeus), infant. Sarawak, Malaysia (Borneo).
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  • Spinomantis elegans, a cave-dwelling frog from high mountains in southeastern Madagascar. Ranomafana National Park, Madagascar.
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  • Owston's Palm Civet (Chrotogale owstoni), a nocturnal carnivore from Laos and Vietnam, is a highly vulnerable species due to illegal poaching. Captive. Endangered Primate Rescue Centre, Cuc Phuong National Park, Vietnam.
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  • Formally described in 2009, this pitcher plant (Nepenthes bokorensis) is endemic to the Kampot province of southern Cambodia where it grows on rocky montane plateaus amid scrub and short forest.
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  • Like some kind of karmic balance, it has often been remarked that the more plain-looking a bird is, the more beautiful its song, whereas those bestowed with gorgeous plumage tend to have simple calls. Research across hundreds of bird species has revealed a negative correlation between these traits, suggesting that there may be a trade-off in visual vs. acoustic signals when it comes to attracting mates. Likely falling at one end of this spectrum, Birds-of-paradise certainly are among the most extravagantly ornamented species in the world, but their highly varied calls often consist of harsh squawks and penetrating whistles, which although extremely loud are anything but melodious. Here, a male Wilson’s BoP (Cicinnurus respublica) raucously advertises his presence above his display ground, though if a female approaches he will fall silent and rely on his feathers for the real game of courtship. Waigeo Island, West Papua, Indonesia.
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  • Scat of Mountain Treeshrew (Tupaia montana) on the peristome of Nepenthes rajah. This will be washed into the pitcher with rain where it becomes a vital source of nutrients for the plant. Sabah, Malaysia (Borneo).
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  • Borneo Necklaced Partridge (Tropicoperdix graydoni), considered by some to be a subspecies of Chestnut-necklaced Partridge (Tropicoperdix charltonii) from mainland Southeast Asia. Sabah, Malaysia (Borneo).
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  • The least known of all of Sulawesi's monkeys, Heck's Macaque occurs on the northwestern portion of the island. Five other endemic species of macaque are distributed in different regions of Sulawesi, occasionally producing hybrids where their ranges overlap. All are diurnal and feed largely on fruits. Central Sulawesi, Indonesia.
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  • Compact clumps of the rare pitcher plant Nepenthes campanulata grow on the vertical walls of a limestone cliff nearly a hundred meters above the canopy of the rainforest. Sarawak, Malaysia (Borneo).
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  • Madagascar’s largest land predator, the Fossa (Cryptoprocta ferox). This receptive female has climbed a tall tree where she waits for a male who is able to join her on one of the upper branches. Reaching her isn’t always an easy task for the males, which are considerably larger and heavier, and they often face fierce competition when attempting to do so. Kirindy Reserve, Madagascar.
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  • Gursky's Spectral Tarsier (Tarsius spectrumgurskyae), juvenile. North Sulawesi, Indonesia.
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  • The Earless Monitor Lizard (Lanthanotus borneensis) is endemic to Borneo where it lives in rocky stream habitats in lowland rainforest. Because of its extreme rarity (it is known from only a few specimens), its life history remains unstudied. It is the only species within its entire family (Lanthanotidae) and was once though to be the missing link between lizards and snakes. Sarawak, Malaysia (Borneo).
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  • The Maroon Langur (Presbytis rubicunda) is endemic to Borneo and nearby Karimata Island, with several subspecies occurring throughout its range. Like other leaf monkeys they are almost completely arboreal and feed on foliage, seeds, and some fruits.  Danum Valley Conservation Area, Sabah, Malaysia.
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  • The Smooth Otter (Lutrogale perspicillata) is the largest otter species in Southeast Asia, reaching up to 1.3 meters in length. Smooth Otters mate for life and hunt cooperatively for fish. Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Pitcher plant (Nepenthes lowii). Sarawak, Malaysia (Borneo).
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  • Two-banded Chameleon (Furcifer balteatus), female. Ranomafana National Park, Madagascar.
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  • Goodman's Mouse Lemur (Microcebus lehilahytsara), feeding on the nectar of night-blooming Dombeya flowers. Andasibe, Madagascar.
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  • A Small-toothed Sportive Lemur (Lepilemur microdon) peers curiously from its tree hole in the montane forest of Ranomafana National Park, Madagascar. Usually emerging only at night, sportive lemurs frequently wake to keep a watchful eye out for intruders into their territory. Not long ago only 8 species of Lepilemur were known from Madagascar, but recent molecular work has resulted in at least 26 distinct species now being recognized, with more sure to follow.
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  • A Bornean Bearded Pig (Sus barbatus) forages for food in a mangrove swamp during low tide.
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  • Lion (Panthera leo). Moremi Game Reserve, Ngamiland, Botswana.
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  • A young Parson's Chameleon (Calumma parsonii) in the rainforest of Masoala National Park. Antsiranana, Madagascar.
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  • At only 12cm in length, the Sulawesi Dwarf Kingfisher (Ceyx fallax) is one of the smallest kingfishers. It is a bird of the forest, feeding on small lizards and insects, and is endemic to Sulawesi and a few offshore islands, although scarce and patchily distributed. Gorontalo, Indonesia.
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  • A large male Bornean Clouded Leopard (Neofelis diardi) is caught on camera trap as he patrols his territory in the rainforest of Danum Valley, Sabah. Borneo's largest cat species, the clouded leopard preys on a variety of smaller wildlife ranging from deer and wild boar to monkeys. Sabah, Malaysia (Borneo).
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  • The critically endangered Greater Bamboo Lemur (Prolemur simus) is restricted to wet rainforests of Madagascar's southeast where it is highly dependent on Cathariostachys bamboo. Ranomafana National Park, Madagascar.
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  • Large Flying Fox (Pteropus vampyrus). Sarawak, Malaysia (Borneo).
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  • Nepenthes macrophylla. This rare montane pitcher plant is endemic only to the summit of Gunung Trusmadi, Malaysia's second highest mountain. Sabah, Malaysia (Borneo).
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  • Mareeba Rock Wallaby (Petrogale mareeba). Queensland, Australia.
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  • Rhinoceros Hornbill (Buceros rhinoceros). Sepilok Forest Reserve, Sabah, Malaysia (Borneo).
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  • Rufous-headed Ground-roller (Atelornis crossleyi). Ranomafana National Park, Madagascar.
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  • A Red-shanked Douc (Pygathrix nemaeus) munches on a handful of young vine leaves it has picked in the forest canopy. Although these endangered monkeys feed on over sixty different species of plants, they select their food carefully and in particular avoid sweet fruits which can upset the balance of bacteria in their guts. Like other leaf-eating colobine monkeys, doucs have large chambered stomachs – making them the only ruminant primates and giving them their characteristic pot-bellied appearance. Son Tra Nature Reserve, Vietnam.
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  • Ryabov's Bug-eyed Frog (Theloderma ryabovi), female. Thua Thien Hue, Vietnam.
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  • Hose’s Civet (Diplogale hosei), one of Borneo’s rarest and least known endemic mammals. This otter-like carnivore is an inhabitant of high elevation mossy forests where it hunts for frogs, fish, and other small animals. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Northeast Bornean Orangutan (Pongo pygmaeus morio), adult male. Sabah, Malaysia (Borneo).
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  • Similar to the Dendrobatid poison frogs, harlequin toads (Atelopus spp.) are also brightly colored, day-active, and highly toxic. They also comprise one of the most endangered groups of amphibians in the neotropics, with many species listed as Critically Endangered and others presumed extinct in the wild. Aside from habitat loss and threats from introduced species, they appear particularly susceptible to the amphibian-killing chytrid fungus, which has completely wiped out many wild populations. This Atelopus spurrelli is endemic to the Chocóan rainforest of coastal Colombia.
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  • A pair of Madagascar Hoopoes (Upupa marginata) perch outside of their cliffside nest burrow. Isalo National Park, Madagascar.
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  • Schlegel's Asity (Philepitta schlegeli), endemic to Madagascar. Tsingy de Bemaraha National Park, Madagascar.
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  • Vogelkop Ringtail (Pseudochirulus schlegeli), one of New Guinea’s least studied possums, being known from only a few specimens. Like other ringtails, they are strictly nocturnal, emerging at night to feed on fruit and leaves. Arfak Mountains, West Papua, Indonesia (New Guinea).
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  • Silvered Leaf Monkey (Trachypithecus cristatus). Sarawak, Malaysia (Borneo).
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  • The Crested Black Macaque (Macaca nigra nigra) is one of four monkey species endemic to Sulawesi.  They are sometimes wrongly referred to as apes because their tail is extremely small and difficult to see.  They travel mostly on the ground in large groups of up to nearly 100 individuals. North Sulawesi, Indonesia.
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  • A newly distinguished species, the Kayan Loris (Nycticebus kayan) differs from other Bornean loris species by a combination of morphological traits. Sarawak, Malaysia (Borneo).
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  • This large male Northwest Bornean Orangutan (Pongo pygmaeus pygmaeus) is just beginning to develop cheek pads, a sign of maturity and placing his age to perhaps between 15 to 20 years. Batang Ai, Sarawak, Malaysia (Borneo).
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  • The Earless Monitor Lizard (Lanthanotus borneensis) is endemic to Borneo where it lives in rocky stream habitats in lowland rainforest. Because of its extreme rarity (it is known from only a few specimens), its life history remains unstudied. It is the only species within its entire family (Lanthanotidae) but it is related to the true monitor lizards (Varanidae). Sarawak, Malaysia (Borneo).
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  • Although shy, the Bearded Pig (Sus barbatus) is abundant in the forests of Borneo, and its presence is usually indicated by diggings and wallows left as they forage for food on the forest floor. Sarawak, Malaysia (Borneo).
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  • The newly distinguished Kayan Loris (Nycticebus kayan) differs from other Bornean loris species by a combination of morphological traits. Sarawak, Malaysia (Borneo).
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  • Red-fronted Brown Lemur (Eulemur rufifrons)
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  • Feared by some, revered by others, the Aye-aye (Daubentonia madagascariensis) is surely one of Madagascar's most bizarre lemurs. Unique among primates, the Aye-aye's front incisors grow continuously like a rodent, leading to the early belief that they were some sort of gigantic nocturnal squirrel. Equally unusual are its long bony middle fingers which are used by the animal to tap tree trunks for the sound of insect grubs inside and then extract them like a fish hook. Unfortunately, Aye-ayes have become gravely endangered from habitat loss and persecution by people: they are often killed on sight due to the superstition that they are an omen of death. Tomasina, Madagascar.
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  • The endangered Bear Monkey (Trachypithecus vetulus monticola) is endemic the mountain forests of Sri Lanka. Central Province, Sri Lanka.
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  • Mountain Gorilla (Gorilla beringei beringei). Volcanoes National Park, Ruhengeri, Rwanda.
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  • This unusual species of pitcher plant (Nepenthes klossii) bears a concealed opening to deceive insect prey. It is extremely rare, only growing in remote high elevaton swampy forests in New Guinea. Papua, Indonesia.
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  • The rare and enigmatic Babirusa (Babyrousa celebensis) occurs only on Sulawesi and a few neighboring islands. Adult males have four conspicuous tusks which are formed from their elongated canine teeth. Gorontalo, Sulawesi, Indonesia.
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  • This large male Northwest Bornean Orangutan (Pongo pygmaeus pygmaeus) is just beginning to develop cheek pads, a sign of maturity and placing his age to perhaps between 15 to 20 years. Batang Ai, Sarawak, Malaysia (Borneo).
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  • A Gursky's Spectral Tarsier (Tarsius spectrumgurskyae) perches outside of its tree hole roost. These tiny nocturnal primates emerge every evening to hunt for insects. Tangkoko Nature Reserve, North Sulawesi, Indonesia.
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  • Geoffroy’s Tamarin (Saguinus geoffroyi). Colón, Panama.
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  • Canopy Slug-eating Snake (Sibon canopy), a new species described in January 2023, endemic to Panama. Panamá Oeste, Panama.
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  • Eastern African Cheetah (Acinonyx jubatus jubatus). Serengeti National Park, Tanzania.
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  • Wide-headed Tree Frog (Leptopelis grandiceps). Udzungwa Mountains, Tanzania.
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  • Silvered Leaf Monkey (Trachypithecus cristatus), rare white morph. Bako National Park, Sarawak, Malaysia (Borneo).
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  • Rufous-collared Kingfisher (Actenoides concretus), female. Sarawak, Malaysia (Borneo).
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  • The bizarre and rare Bornean Bristlehead (Pityriasis gymnocephala) is one of the most enigmatic birds of Borneo and is highly sought after by birdwatchers. It is the only member of its family and its nearest relations to other birds has been highly debated. Bristleheads are found in small nomadic flocks traveling in the middle to upper canopy where they feed on insects. Sabah, Malaysia (Borneo).
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  • A large male O'Shaughnessy's Chameleon (Calumma oshaughnessyi) stalks up a forest liana, keeping an eye out for both prey and predator alike. With 360-degree vision, remarkable image acuity, and monocular depth perception, chameleons can detect insects that might be well out of reach, judge their distance accurately, and even distinguishing them from their surroundings when the insect is completely still. Ranomafana National Park, Madagascar.
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  • Blooming high in a moss-laden tree of Ranomafana’s montane forest, this epiphytic orchid (Aerangis fastuosa) exhales an alluring perfume at night to attract its desired pollinator: hawk moths. Madagascar has many such night-active orchids, most of which sport long tapered nectaries that can only be reached by the long tongues of their associated moths. With over a thousand species, the orchids of Madagascar are exceptionally diverse. Ranomafana National Park, Madagascar.
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  • Kayan Loris (Nycticebus kayan), juvenile. Gunung Mulu National Park, Sarawak, Malaysia (Borneo).
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  • A Gursky's Spectral Tarsier (Tarsius spectrumgurskyae) perches outside of its tree hole roost. These tiny nocturnal primates emerge every evening to hunt for insects. Tangkoko Nature Reserve, North Sulawesi, Indonesia.
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  • Pitcher plant (Nepenthes bicalcarata). Normally occuring only in shady peat swamp forests, this species occasionally grows in open heath-like peat bogs. It is endemic to Borneo. Sarawak, Malaysia.
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  • The Sumatran Surili (Presbytis melalophos) is endemic to the island of Sumatra.  Like other leaf monkeys, this species is primarily folivorous, but also consumes flowers and fruits. Kerinci Seblat National Park, West Sumatra, Indonesia.
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