Chien C. Lee

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  • In the coastal forests of northern Borneo, a young Silvery Lutung (Trachypithecus cristatus) is groomed by its mother. Like many leaf monkey species, these highly social primates display brightly colored fur when born that gradually fades after several months into their adult coloration. The exact reason for this adaption is uncertain, but it may help in protection: members of the group can easily distinguish their vulnerable young at a glance, or locate them quickly in the dense foliage of the rainforest canopy. Another interesting theory is that certain predators such as Clouded Leopards, are red/green colorblind, so an orangish pelt may actually be an effective camouflage. Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • A captive Bear Cuscus (Ailurops ursinus) peers out of a small wooden cage behind a village in Central Sulawesi. Listed as Vulnerable by the IUCN Red List, the Bear Cuscus is endemic to Sulawesi and a few offshore islands where it is restricted to undisturbed rainforest. Central Sulawesi, Indonesia.
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  • Nepenthes distillatoria, a Sri Lankan endemic and the only pitcher plant species occurring on the island.
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  • Nycticebus menagensis. One of four species of Slow Loris found in Borneo. All Slow Loris are nocturnal arboreal primates which feed on small animals, insects and soft fruits. Sarawak, Malaysia (Borneo).
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  • Endemic to the mountains of central Sulawesi, the stream-breeding Loka Flying Frog (Rhacophorus monticola) is highly variable in coloration. South Sulawesi, Indonesia.
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  • Pitcher plant (Nepenthes edwardsiana). Sabah, Malaysia (Borneo).
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  • Lion (Panthera leo). Moremi Game Reserve, Ngamiland, Botswana.
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  • Where their ranges overlap, Heck's Macaque (Macaca hecki) and Tonkean Macaque (Macaca tonkeana) form a natural hybrid population, with all these individuals showing intermediate characteristics between the two species. Central Sulawesi, Indonesia.
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  • Bulwer's Pheasant (Lophura bulweri), male. Sabah, Malaysia (Borneo).
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  • Yellow Mantella (Mantella crocea). Andasibe, Madagascar.
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  • With its rough skin texture and mottled green coloration, it’s easy to see how the Mossy Rain Frog (Pristimantis museosus) is well equipped to blend in with the damp forests in which it dwells. This frog is endemic to the mountains of central Panama, a region which has seen drastic declines in many amphibians over the past few decades due to the introduction of the virulent chytrid fungus. Like other species of Pristimantis, the females of P. museosus lay large eggs that do not require water – the tadpoles transform into froglets entirely within the egg itself. This adaptation may have provided this species with some protection against chytrid, since the fungus most easily transfers among those species which breed in bodies of water. Nevertheless, this frog remains highly threatened and has disappeared from multiple sites across its range. Panamá Oeste, Panama.
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  • Usambara Eyelash Viper (Atheris ceratophora). Udzungwa Mountains, Tanzania.
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  • The Palawan Peacock Pheasant (Polyplectron napoleonis) is a shy ground-dwelling bird endemic to the rainforests the island of Palawan, Philippines.
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  • Madagascar Plover (Charadrius thoracicus). Toliara, Madagascar.
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  • Hose’s Civet (Diplogale hosei), one of Borneo’s rarest and least known endemic mammals. This otter-like carnivore is an inhabitant of high elevation mossy forests where it hunts for frogs, fish, and other small animals. Sarawak, Malaysia (Borneo).
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  • Endemic to the Bird's Head Peninsula of western New Guinea, the Vogelkop Tree Kangaroo (Dendrolagus ursinus) has become increasingly threatened by habitat loss and hunting. West Papua, Indonesia (New Guinea).
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  • The Diabolical Nightjar (Eurostopodus diabolicus) produces such an eerie call that local people in Sulawesi hold the superstitious belief that it is a demonic entity, hence its name. The life history of this rare bird is very poorly known as until recently only a single specimen had ever been found. Like other nightjars it rests by day on the ground, using its superb camouflage to escape detection from predators. Central Sulawesi, Indonesia.
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  • Pitcher plant (Nepenthes lowii). Sarawak, Malaysia (Borneo).
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  • Goodman's Mouse Lemur (Microcebus lehilahytsara), feeding on the nectar of night-blooming Dombeya flowers. Andasibe, Madagascar.
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  • Niah Cave Gecko (Cyrtodactylus cavernicolus). Sarawak, Malaysia (Borneo).
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  • Rhinoceros Hornbill (Buceros rhinoceros). Sepilok Forest Reserve, Sabah, Malaysia (Borneo).
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  • Pitcher plant (Nepenthes campanulata), a limestone lithophytic carnivorous plant, here growing on a cliff 250m above the rainforest canopy. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • A Maroon Langur (Presbytis rubicunda) pauses for a break while munching on his chosen leaf. Endemic to Borneo and a few offshore islands, these leaf monkeys feed on a wide range of foliage and unripe fruits, traveling in groups of up to a dozen individuals. One theory regarding their distinctive bright orange coloration is that this may help to confuse some predators into mistaking them for a much larger and stronger Orangutan, and thus deter attacks. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Heck's Macaque (Macaca hecki) x Tonkean Macaque (Macaca tonkeana), natural hybrid population. Central Sulawesi, Indonesia.
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  • Sri Lanka Rock Frog (Nannophrys ceylonensis). Sinharaja National Park, Sri Lanka.
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  • The bright orange coloration of this infant Silvered Langur (Trachypithecus cristatus) helps them to be seen and hence guarded by the entire troop. They will gradually lose this coloration after 3 months of age. Bako National Park, Sarawak, Malaysia (Borneo).
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  • A pair of Madagascar Hoopoes (Upupa marginata) perch outside of their cliffside nest burrow. Isalo National Park, Madagascar.
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  • Compact clumps of the rare pitcher plant Nepenthes campanulata grow on the vertical walls of a limestone cliff nearly a hundred meters above the canopy of the rainforest. Sarawak, Malaysia (Borneo).
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  • Madagascar’s largest land predator, the Fossa (Cryptoprocta ferox). This receptive female has climbed a tall tree where she waits for a male who is able to join her on one of the upper branches. Reaching her isn’t always an easy task for the males, which are considerably larger and heavier, and they often face fierce competition when attempting to do so. Kirindy Reserve, Madagascar.
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  • Vogelkop Ringtail (Pseudochirulus schlegeli), one of New Guinea’s least studied possums, being known from only a few specimens. Like other ringtails, they are strictly nocturnal, emerging at night to feed on fruit and leaves. Arfak Mountains, West Papua, Indonesia (New Guinea).
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  • Gursky's Spectral Tarsier (Tarsius spectrumgurskyae), juvenile. North Sulawesi, Indonesia.
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  • Heck's Macaque (Macaca hecki) x Tonkean Macaque (Macaca tonkeana), natural hybrid population. Central Sulawesi, Indonesia.
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  • Silvered Leaf Monkey (Trachypithecus cristatus). Sarawak, Malaysia (Borneo).
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  • The Maroon Langur (Presbytis rubicunda) is endemic to Borneo and nearby Karimata Island, with several subspecies occurring throughout its range. Like other leaf monkeys they are almost completely arboreal and feed on foliage, seeds, and some fruits.  Danum Valley Conservation Area, Sabah, Malaysia.
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  • The Smooth Otter (Lutrogale perspicillata) is the largest otter species in Southeast Asia, reaching up to 1.3 meters in length. Smooth Otters mate for life and hunt cooperatively for fish. Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Although shy, the Bearded Pig (Sus barbatus) is abundant in the forests of Borneo, and its presence is usually indicated by diggings and wallows left as they forage for food on the forest floor. Sarawak, Malaysia (Borneo).
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  • A Bornean Bearded Pig (Sus barbatus) forages for food in a mangrove swamp during low tide.
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  • The newly distinguished Kayan Loris (Nycticebus kayan) differs from other Bornean loris species by a combination of morphological traits. Sarawak, Malaysia (Borneo).
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  • Rather than being carnivorous, this unusual pitcher plant (Nepenthes lowii) derives its nutrition from the droppings of the Mountain Treeshrew (Tupaia montana). The animals are attracted to the plant's copious nectar secretions, and inevitably leave their scat in the pitchers which are designed like a natural toilet receptacle. Sarawak, Malaysia (Borneo).
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  • The rare and enigmatic Babirusa (Babyrousa celebensis) occurs only on Sulawesi and a few neighboring islands. Adult males have four conspicuous tusks which are formed from their elongated canine teeth. Gorontalo, Sulawesi, Indonesia.
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  • A Gursky's Spectral Tarsier (Tarsius spectrumgurskyae) perches outside of its tree hole roost. These tiny nocturnal primates emerge every evening to hunt for insects. Tangkoko Nature Reserve, North Sulawesi, Indonesia.
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  • Variable Cross Frog (Oreophryne variabilis). Widespread in Sulawesi, the loud calls of this tiny bush frog are a common sound in cool and wet mossy montane forests. South Sulawesi, Indonesia.
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  • Eastern African Cheetah (Acinonyx jubatus jubatus). Serengeti National Park, Tanzania.
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  • Gaboon Viper (Bitis gabonica). Udzungwa Mountains, Tanzania.
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  • Wide-headed Tree Frog (Leptopelis grandiceps). Udzungwa Mountains, Tanzania.
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  • Kerangas Bush Frog (Philautus kerangae). Sarawak, Malaysia (Borneo).
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  • A large male O'Shaughnessy's Chameleon (Calumma oshaughnessyi) stalks up a forest liana, keeping an eye out for both prey and predator alike. With 360-degree vision, remarkable image acuity, and monocular depth perception, chameleons can detect insects that might be well out of reach, judge their distance accurately, and even distinguishing them from their surroundings when the insect is completely still. Ranomafana National Park, Madagascar.
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  • A Horsfield's Tarsier (Cephalopachus bancanus) on the prowl for food in the rainforest of western Borneo. These tiny primates are exclusively carnivorous, hunting insects, small lizards, and even snakes by leaping deftly through the thick forest understory and snatching prey with their long nimble fingers. Unlike most other nocturnal mammals which possess a reflective layer behind their retina to help with night vision, tarsiers instead compensate by having enormous eyes. In fact, this species sports the largest eyes in relation to its body size of any mammal in the world - these are locked in place and the animal is only able to shift its vision by rotating its head. Sarawak, Malaysia (Borneo).
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  • Pitcher plant (Nepenthes sibuyanensis). This large pitcher plant is endemic to the ultramafic Mount Guiting-guiting on Sibuyan Island, Philippines.
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  • Kayan Loris (Nycticebus kayan), juvenile. Sarawak, Malaysia (Borneo).
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  • Pitcher plant (Nepenthes bicalcarata). Normally occuring only in shady peat swamp forests, this species occasionally grows in open heath-like peat bogs. It is endemic to Borneo. Sarawak, Malaysia.
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  • Red-fronted Brown Lemur (Eulemur rufifrons). Ranomafana National Park, Madagascar.
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  • Lesser Adjutant (Leptoptilos javanicus). Yala National Park, Sri Lanka.
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  • Brown Mouse Lemur (Microcebus rufus). Ranomafana National Park, Madagascar.
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  • Sulawesi Babirusa (Babyrousa celebensis). Gorontalo, Sulawesi.
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  • Silvered Leaf Monkey (Trachypithecus cristatus). Sarawak, Malaysia (Borneo).
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  • Blue-headed Pitta (Hydrornis baudii), male. Sabah, Malaysia (Borneo).
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  • Endemic to the Bird's Head Peninsula of western New Guinea, the Vogelkop Tree Kangaroo (Dendrolagus ursinus) has become increasingly threatened by habitat loss and hunting. West Papua, Indonesia (New Guinea).
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  • Once considered one of the three most elusive birds in the world, the Long-tailed Ground Roller (Uratelornis chimaera) is a prized sighting for any birdwatcher. The entire family of Ground Rollers (Brachypteraciidae, 5 species) is endemic to Madagascr, but while most are inhabitants of rainforest, this is the only species found in in the arid spiny desert of the island’s extreme southwest. Even within this region it has as very restricted distribution and occurs at low densities, with only a few birds per square kilometer. Adapted for running swiftly on the ground with their powerful legs, these birds hunt for insects and small animals in the sand and leaf-litter. Toliara, Madagascar.
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  • Pitcher plant (Nepenthes campanulata). Sarawak, Malaysia (Borneo).
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  • One of the most beautiful of all pitcher plants, Nepenthes edwardsiana produces enormous cylindrical pitchers which are strikingly colored. It is endemic to the Mount Kinabalu region in northern Borneo.
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  • The Maroon Langur (Presbytis rubicunda) is endemic to Borneo and nearby Karimata Island, with several subspecies occurring throughout its range. Like other leaf monkeys they are almost completely arboreal and feed on foliage, seeds, and some fruits. Sabah, Malaysia.
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  • Marojejy Mantella (Mantella manery). Marojejy National Park, Madagascar.
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  • A newly distinguished species, the Kayan Loris (Nycticebus kayan) differs from other Bornean loris species by a combination of morphological traits. Sarawak, Malaysia (Borneo).
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  • An adult male Silvered Leaf Monkey (Trachypithecus cristatus) keeps a watchful eye out for any threats to his harem of females. Sarawak, Malaysia (Borneo).
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  • White-fronted Brown Lemur (Eulemur albifrons), male. Marojejy National Park, Antsiranana, Madagascar.
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  • A Northern Bamboo Lemur (Hapalemur occidentalis) in the foothills of Marojejy National Park, one of at least five species of Hapalemur distributed throughout the forests of Madagascar. True to their name, these small lemurs subsist on a diet that consists largely of young bamboo leaves, a remarkable feat considering the high cyanide content found in the plant’s tissues. It is still not completely understood exactly how these lemurs are able to survive on a diet so tainted with cyanide that it would kill animals much larger than themselves. Antsiranana, Madagascar.
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  • Red-fronted Brown Lemur (Eulemur rufifrons)
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  • Leopard (Panthera pardus), juvenile. Ngamiland, Botswana.
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  • Giraffe (Giraffa camelopardalis giraffa). Mpumalanga, South Africa.
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  • The Maroon Langur (Presbytis rubicunda), also known as the Red Leaf Monkey, is endemic to Borneo and nearby Karimata Island, with several subspecies occurring throughout its range. Like other leaf monkeys they are almost completely arboreal and feed on foliage, seeds, and some fruits. Sabah, Malaysia.
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  • Knobbed Hornbill (Rhyticeros cassidix), male perched outside of tree hole nest where he passes food to the female enclosed within. North Sulawesi, Indonesia.
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  • Gaboon Viper (Bitis gabonica). Udzungwa Mountains, Tanzania.
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  • Gaboon Viper (Bitis gabonica). Udzungwa Mountains, Tanzania.
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  • The fox-like Fanaloka (Fossa fossana) is sometimes referred to as a “Malagasy Civet” but it actually belongs to the endemic Madagascan family Eupleridae. Furtive and nocturnal, it is the second-largest native carnivore on the island, preying on small animals that it hunts on the forest floor. Ranomafana National Park, Madagascar.
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  • Silvered Leaf Monkey (Trachypithecus cristatus), rare white morph. Bako National Park, Sarawak, Malaysia (Borneo).
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  • Sunda Clouded Leopard (Neofelis diardi). Sarawak, Malaysia (Borneo).
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  • Dung-eating pitcher plant (Nepenthes lowii), endemic to Borneo. Pulong Tau National Park, Sarawak, Malaysia.
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  • A Gursky's Spectral Tarsier (Tarsius spectrumgurskyae) perches outside of its tree hole roost. These tiny nocturnal primates emerge every evening to hunt for insects. Tangkoko Nature Reserve, North Sulawesi, Indonesia.
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  • Maroon Langur (Presbytis rubicunda) feeding on leaves of Mikania micrantha, an invasive vine. Danum valley, Sabah, Malaysia (Borneo).
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  • In the dim understory of the forest floor the color of this poison dart frog (Oophaga solanensis) seem almost too fantastic to be real. Like other dendrobatid frogs, these colors serve as a warning signal of their toxicity. This species is endemic to the coastal rainforests of Colombia. Choco, Colombia.
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  • The Smooth Otter (Lutrogale perspicillata) is the largest otter species in Southeast Asia, reaching up to 1.3 meters in length. Smooth Otters mate for life and hunt cooperatively for fish. Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • After several months of feeding on Aristolochia leaves, the mature caterpillar of this large birdwing butterfly (Troides andromache) prepares for its final moult into the pupal stage when metamorphosis will transform it into an adult. Sabah, Malaysia (Borneo).
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  • The Western Tarsier (Cephalopachus bancanus borneanus), also known as Horsfield's Tarsier, is a small nocturnal primate occurring in the lowland forests of Sumatra and Borneo. They are carnivorous and feed mainly on insects, but occasionally take birds and bats. Sarawak, Malaysia (Borneo).
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  • Nepenthes argentii. With traps usually less than 3 cm in height this species is perhaps the smallest Nepenthes pitcher plant in the world. It is endemic to Mt. Guiting-guiting on Sibuyan Island, where it grows among stunted alpine vegetation. Romblon, Philippines.
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  • A non-climbing species, this pitcher plant (Nepenthes campanulata) grows only on the vertical faces of tall limestone cliffs in Borneo. It is so difficult to observe in habitat that it was once thought to be extinct. Sarawak, Malaysia (Borneo).
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  • Pitcher plant (Nepenthes northiana). Sarawak, Malaysia (Borneo).
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  • Rufous Mouse Lemur (Microcebus rufus), feeding on katydid. Ranomafana National Park, Madagascar.
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  • Blooming high in a moss-laden tree of Ranomafana’s montane forest, this epiphytic orchid (Aerangis fastuosa) exhales an alluring perfume at night to attract its desired pollinator: hawk moths. Madagascar has many such night-active orchids, most of which sport long tapered nectaries that can only be reached by the long tongues of their associated moths. With over a thousand species, the orchids of Madagascar are exceptionally diverse. Ranomafana National Park, Madagascar.
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  • Yellow-bellied Sunbird-asity (Neodrepanis hypoxantha), male in breeding plumage. Despite their appearance, these birds are closely related to Asian broadbills; their slender beaks representing an example of convergence with sunbirds due to their nectar-feeding habits. Ranomafana National Park, Madagascar.
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  • Brown Mouse Lemur (Microcebus rufus). Ranomafana National Park, Madagascar.
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  • The handsomely-proportioned Short-legged Ground-roller (Brachypteracias leptosomus) belongs to yet another of Madagascar’s endemic bird families, Brachypteraciidae. Patchily distributed in the rainforests of eastern Madagascar, it is a seldom-seen species owing to its habit of perching completely still and quiet for long periods of time in the lower canopy. They feed on large insects and other invertebrates. Ranomafana National Park, Madagascar.
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  • Yellow-bellied Sunbird-asity (Neodrepanis hypoxantha), male in breeding plumage. Despite their appearance, these birds are closely related to Asian broadbills; their slender beaks representing an example of convergence with sunbirds due to their nectar-feeding habits. Ranomafana National Park, Madagascar.
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  • Bornean Crested Fireback (Lophura ignita), male. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Bornean Sun Bear (Helarctos malayanus euryspilus). Bornean Sun Bear Conservation Centre, Sabah, Malaysia (Borneo).
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  • The Waigeo Spotted Cuscus (Spilocuscus papuensis) is endemic to a single island off the northwestern coast of West Papua. Closely related to possums, this is a pouched marsupial, spending its entire life in the trees where it feeds on leaves and occasional fruits.
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  • This unusual pitcher plant (Nepenthes lowii) derives its nutrition from the droppings of the Mountain Treeshrew (Tupaia montana). The animals are attracted to the plant's copious nectar secretions, and inevitably leave their scat in the pitchers which are designed like a natural toilet receptacle. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • The bizarre gourd-shaped pitchers of this pitcher plant (Nepenthes lowii) are an unforgettable sight in the mossy forests of northern Borneo. It is has been shown that these function to trap animal droppings which serve as fertilizer for the plant. Sarawak, Malaysia (Borneo).
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  • Pitcher plant (Nepenthes pitopangii). Central Sulawesi, Indonesia.
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  • Like other langurs, Silvered Leaf Monkeys (Trachypithecus cristatus) are highly social and travel in cohesive groups numbering up to 40 or more individuals. They have a specialized diet, feeding almost exclusively on high-protein leaves. Sarawak, Malaysia (Borneo).
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