Chien C. Lee

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  • Plain-pouched Hornbill (Rhyticeros subruficollis), male at nest. Huai Kha Khaeng Wildlife Sanctuary, Thailand.
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  • Crested Serpent Eagle (Spilornis cheela). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Brown-tailed Racer (Gonyosoma oxycephalum). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • The Smooth Otter (Lutrogale perspicillata) is the largest otter species in Southeast Asia, reaching up to 1.3 meters in length. Smooth Otters mate for life and hunt cooperatively for fish. Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Estuarine Crocodile (Crocodylus porosus). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Estuarine Crocodile (Crocodylus porosus). Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • In the coastal forests of northern Borneo, a young Silvery Lutung (Trachypithecus cristatus) is groomed by its mother. Like many leaf monkey species, these highly social primates display brightly colored fur when born that gradually fades after several months into their adult coloration. The exact reason for this adaption is uncertain, but it may help in protection: members of the group can easily distinguish their vulnerable young at a glance, or locate them quickly in the dense foliage of the rainforest canopy. Another interesting theory is that certain predators such as Clouded Leopards, are red/green colorblind, so an orangish pelt may actually be an effective camouflage. Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Borneo Pygmy Elephant (Elephas maximus borneensis). Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Sunda Clouded Leopard (Neofelis diardi), female. Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Red-bearded Bee-eater (Nyctyornis amictus), male. Tabin Wildlife Reserve, Sabah, Malaysia (Borneo).
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  • Bronzeback (Dendrelaphis terrificus). Gorontalo, Sulawesi, Indonesia.
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  • Smith's Litter Frog (Leptobrachium smithi). Krabi, Thailand.
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  • The largest terrestrial and river predator in the world, the Estuarine Crocodile (Crocodylus porosus) can reach lengths of over 6 m and consumes a wide range of animal prey. They are commonly found in mangroves, lagoons, and large rivers along the coastline of Borneo.  Kinabatangan River, Sabah, Malaysia (Borneo).
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  • The Asian Eyebrow-ridge Toad (Duttaphrynus melanostictus) is a common species widespread in South and Southeast Asia.  It has a distinctive large parotoid glands behind the head, which are known to secrete a toxic substance when the toad is disturbed.
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  • Endemic to Sulawesi and the Sula Islands, the Sulawesi Black Pigeon (Turacoena manadensis) inhabits coastal, lowland, and hill forests. Gorontalo, Indonesia.
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  • A large group of Proboscis Monkeys (Nasalis larvatus) prepares for sleep in a tree overhanging a river. Sabah, Malaysia (Borneo).
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  • Closeup portrait of a Tokay Gecko (Gekko gecko). Uthai Thani, Thailand.
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  • Terciopelo (Bothrops asper). Cartago, Costa Rica.
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  • Smith's Green-eyed Gecko (Gekko smithii). Kinabatangan River, Sabah, Malaysia (Borneo).
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  • Harlequin Tree Frog (Rhacophorus pardalis). Sabah, Malaysia (Borneo).
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  • A male Black-naped Monarchs (Hypothymis azurea) at his nest. In these flycatchers both male and female will share equal care in tending to the nestlings. Krabi, Thailand.
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  • An adult Common Butterfly Lizard (Leiolepis belliana) keeps a watchful eye out for predators whilst her young remain at the entrance of the nest burrow.  Nine species of butterfly lizards are known from the drier sandy regions of Southeast Asia, but their life-history is poorly documented. Uthai Thani, Thailand.
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  • A crystal-clear stream flows through the forest at Khao Nor Chuchi Wildlife Sanctuary.  These spring-fed waters which emerge from limestone rock are highly alkaline and contain high amounts of dissolved calcium carbonate. Krabi, Thailand.
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  • Mixed Deciduous Forest during the beginning of the rainy season in Huai Kha Khaeng Wildlife Sanctuary.
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  • With his crop full of food and water, a male Plain-pouched Hornbill (Rhyticeros subruficollis) returns to his nest in the cavity of a tree wherein the female waits sealed inside. During the nesting period, which can last several months, his mate will be completely dependent on his regular visits to provide her with everything she needs while she incubates the young and raises the chicks. Unfortunately, his duties on this day have been complicated by the annoying attentions of a Greater Racket-tailed Drongo (Dicrurus paradiseus) which has perhaps also been nesting nearby. Although much smaller than the hornbill, drongos are known for their bold and aggressive behavior, especially towards other birds that could be potential predators of their nests. Huai Kha Khaeng Wildlife Sanctuary, Thailand.
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  • Dense bamboo forest along edges of Huai Kha Khaeng River, consisting mostly of huge stands of Giant Thorny Bamboo (Bambusa bambos). Huai Kha Khaeng Wildlife Sanctuary, Thailand.
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  • Proboscis Monkey (Nasalis larvatus). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Although sometimes weighing over 9 kg, Indri (Indri indri) can make impressive leaps from tree to tree with their powerful long hind legs. Largest of the living lemurs, this critically-endangered species is an icon for the conservation of Madagascar's beleaguered wildlife, and its wailing song never fails to enchant visitors to their forest realm. Occuring only in the tropical rainforests on the island's eastern side, the Indri's natural range has dwindled due to widespread deforestation and hunting, despite their traditionally sacred status. Tomasina, Madagascar.
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  • A large male Bornean Clouded Leopard (Neofelis diardi) is caught on camera trap as he patrols his territory in the rainforest of Danum Valley, Sabah. Borneo's largest cat species, the clouded leopard preys on a variety of smaller wildlife ranging from deer and wild boar to monkeys. Sabah, Malaysia (Borneo).
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  • Without seeing it move, you could almost mistake the Golden Mantella (Mantella aurantiaca) for a bright orange plastic toy. This is one of Madagascar’s most endangered amphibians and is an icon for conservation of the island’s threatened wildlife. Efforts targeted at protecting this frog’s habitat, coupled with ex-situ breeding programs and reintroduction have helped to protect it from extinction in the wild, but it remains highly threatened and is still known to exist at only two small isolated patches of rainforest. Andasibe, Madagascar.
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  • Without seeing it move, you could almost mistake the Golden Mantella (Mantella aurantiaca) for a bright orange plastic toy. This is one of Madagascar’s most endangered amphibians and is an icon for conservation of the island’s threatened wildlife. Efforts targeted at protecting this frog’s habitat, coupled with ex-situ breeding programs and reintroduction have helped to protect it from extinction in the wild, but it remains highly threatened and is still known to exist at only two small isolated patches of rainforest. Andasibe, Madagascar.
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  • The Smooth Otter (Lutrogale perspicillata) is the largest otter species in Southeast Asia, reaching up to 1.3 meters in length. Smooth Otters mate for life and hunt cooperatively for fish. Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Lichen Huntsman Spider (Heteropoda boiei) with parasitic biting midges (Ceratopogonidae). Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Sumatran Two-tailed Spider (Hersilia sumatrana). Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Ground beetle (Chlaenius sabahensis). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Some of the most noticeable spiders of the Southeast Asian tropical rainforests are the huntsmans (family Sparassidae); not only do they reach impressive sizes and are often colorful but because they don’t use webs for their prey capture they can easily be seen hunting in understory vegetation at night. Some, such as this Lichen Huntsman (Pandercetes sp.) are harder to find than most due to their cryptic camouflage. This female specimen rests atop her equally camouflaged egg sac which has been neatly woven into the fold of a dead leaf. Tabin Wildlife Reserve, Sabah, Malaysia (Borneo).
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  • Although usually considered a terrestrial species, the Striped Kukri Snake (Oligodon octolineatus) is an excellent climber and can occasionally be seen climbing tree trunks. Sarawak, Malaysia (Borneo).
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  • Bornean Orangutan (Pongo pygmaeus), infant. Sarawak, Malaysia (Borneo).
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  • A female Bornean Orang-utan (Pongo pygmaeus) with her young infant. Semengoh Forest Reserve, Sarawak, Malaysia (Borneo).
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  • Red-crowned Barbet (Psilopogon rafflesii), a frugivorous bird from the rainforests of Southeast Asia. Sarawak, Malaysia (Borneo).
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  • Hidden in the dense leaf litter of the rainforest floor, an Amazonian Horned Frog (Ceratophrys cornuta) lies in wait for its next meal. With a mouth wider than the length of its body, and a voracious appetite to boot, these frogs can consume prey as large as small reptiles and rodents. This species is widely distributed in the Amazon Basin, but nowhere particularly abundant. Yasuní National Park, Ecuador.
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  • Bornean Opposite-fingered Tree Frog (Feihyla inexpectata), first photographic record for this species in Sarawak. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Sun Glass Frog (Hyalinobatrachium aueroguttatum), male hiding on the underside of a leaf. The pattern on the back of this species mimics a cluster of the frog's eggs, which help to deter attacks by predatory wasps. Esmeraldas, Ecuador.
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  • Sri Lankan Grey Langur (Semnopithecus priam thersites). Yala National Park, Sri Lanka.
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  • Nycticebus menagensis. One of four species of Slow Loris found in Borneo. All Slow Loris are nocturnal arboreal primates which feed on small animals, insects and soft fruits. Sarawak, Malaysia (Borneo).
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  • Malayan Flat-shelled Turtle (Notochelys platynota). Sarawak, Malaysia (Borneo).
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  • Sabah Bamboo Pitviper (Trimeresurus sabahi). Sarawak, Malaysia (Borneo).
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  • Without a single sighting in nearly 90 years, the Borneo Rainbow Toad (Ansonia latidisca) was considered possibly extinct and listed by Conservation International as one of the "10 Most Wanted Amphibians". The species was rediscovered in 2011 by a team of herpetologists from the University of Malaysia Sarawak, although it remains extremely rare and little is known of its ecology or behavior. This brings a glimmer of hope in a time of global mass amphibian extinctions. Sarawak, Malaysia (Borneo).
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  • Flat-nosed Pitviper (Craspedocephalus puniceus). West Java, Indonesia.
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  • The Black-bearded Tomb Bat (Taphozous melanopogon) roosts in caves and large rock crevices, emerging at night to feed on insects high above the forest canopy. It is found throughout much of mainland Southeast Asia as well as Borneo, Sumatra, and Java. It's echolocation calls while flying are audible as high pitched clicks. Sarawak, Malaysia (Borneo).
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  • When threatened, the Knob-tailed Gecko (Nephrurus levis) raises its body off the ground to appear larger in size. Cape Range National Park, Western Australia.
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  • A male Whitehead's Broadbill (Calyptomena whiteheadi) returns to its nest where it passes food to the hungry chick. Both male and female care for the nestlings, providing insects and fruit, as well as removing fecal pellets. This scarcely-seen species is the largest of all the green broadbills (reaching 27cm in length) and is endemic to the montane forests of Borneo. Sabah, Malaysia.
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  • Threatened by habitat loss and hunting pressure, the Crested Black Macaque (Macaca nigra nigra) is now listed as Critically Endangered. This species lives only in forested areas at the northernmost tip of the island of Sulawesi. North Sulawesi, Indonesia.
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  • Endemic to the mountains of central Sulawesi, the stream-breeding Loka Flying Frog (Rhacophorus monticola) is highly variable in coloration. South Sulawesi, Indonesia.
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  • This Collared Nightjar (Gactornis enarratus), a Madagascar endemic, hides itself among the leaf litter of the forest floor while it sleeps. Placed in its own genus, this species is distinct from all other nightjars (family Caprimulgidae) by its plumage, eggs, and behavior. Mantadia National Park, Madagascar.
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  • Polynesian Rat (Rattus exulans). Sarawak, Malaysia (Borneo).
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  • White-spotted Cat Snake (Boiga drapiezii), green morph. Sarawak, Malaysia (Borneo).
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  • Black-headed Cat Snake (Boiga nigriceps). Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Sarayacu Tree Frog (Dendropsophus parviceps). Orellana, Ecuador.
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  • The Rufous Piculet (Sasia abnormis) is the smallest woodpecker in Borneo.  Here a bird is making its territorial call by drumming on a spiny rattan petiole. Sabah, Malaysia (Borneo).
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  • The Palawan Peacock Pheasant (Polyplectron napoleonis) is a shy ground-dwelling bird endemic to the rainforests the island of Palawan, Philippines.
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  • Spotted Cuscus (Spilocuscus maculatus chrysorrhous), juvenile. Papua, Indonesia (New Guinea).
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  • Guatemalan Yellow Grosbeak (Pheucticus chrysopeplus aurantiacus), vocalising. El Triunfo Biosphere Reserve, Chiapas, Mexico.
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  • Long-tailed Sylph (Aglaiocercus kingii). Papallacta, Ecuador.
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  • Owston's Palm Civet (Chrotogale owstoni), a nocturnal carnivore from Laos and Vietnam, is a highly vulnerable species due to illegal poaching. Captive. Endangered Primate Rescue Centre, Cuc Phuong National Park, Vietnam.
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  • Madagascar Plover (Charadrius thoracicus). Toliara, Madagascar.
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  • Spectacled Flying Fox (Pteropus conspicillatus). Waigeo Island, West Papua, Indonesia.
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  • Lesser Bare-backed fruit Bat (Dobsonia minor), feeding on fig. Nimbokrang, Papua, Indonesia (New Guinea).
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  • Larut Bush Frog (Philautus larutensis), male. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Rough-sided Frog (Pulchrana glandulosa). Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Ivohimanita Madagascar Frog (Mantidactylus majori), male guarding eggs. Ranomafana National Park, Madagascar.
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  • Andranolava Reed frog (Heterixalus luteostriatus). Tsaranoro Valley, Madagascar.
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  • Tree frog (Boophis tasymena), male. Ranomafana National Park, Madagascar.
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  • Few creatures possess a mastery of camouflage the way Madagascar’s leaf-tailed geckos do. This Mossy Leaf-tailed Gecko (Uroplatus sikorae) sleeps by day on a rainforest liana, its outline obscured by frilled dermal flaps that line its body. By always resting in a downward-facing position they may be able to quickly catch unsuspecting insects that climb up from the ground and also conceal their reflective lidless eyes from predators lurking above. Ranomafana National Park, Madagascar.
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  • Sumatran Pit Viper (Trimeresurus sumatranus). Sarawak, Malaysia (Borneo).
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  • Boulenger's Digging Frog (Plethodontohyla inguinalis). Ranomafana National Park, Madagascar.
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  • Smallest of all the anteaters, the Silky Anteater (Cyclopes didactylus) is seldom seen because it spends much of its life high in the rainforest canopy, often curled up as an indistinguishable ball of fluff. They have no teeth and can only defend themselves by means of their razor sharp sickle-like fore-claws, which are usually used for tearing open ant nests. Females bear a single youngster at a time, which is carried on their back until large enough to feed on its own. Esmeraldas, Ecuador.
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  • Cloudy Snail-eating Snake (Sibon nebulatus). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • Hump-nosed lizard photo (Lyriocephalus scutatus), male. Sinharaja National Park, Southern Province, Sri Lanka.
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  • Matang Narrow-mouthed Frog (Microhyla nepenthicola), in amplexus.  This is the smallest frog in the Old World and it rears its young exclusively in the fluid of pitcher plants. Sarawak, Malaysia (Borneo).
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  • Wilson's Bird-of-paradise (Diphyllodes respublica), male courting female at his display ground. West Papua, Indonesia (New Guinea)
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  • Brehm's Tiger-parrot (Psittacella brehmii). West Papua, Indonesia (New Guinea).
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  • Endemic to the mountains of northern Borneo, the Kinabalu Horned Frog (Pelobatrachus baluensis) breeds only in clear rocky streams with cold water. By day they lie camouflaged among the leaf litter on the forest floor, but they emerge at night to call for mates. Sabah, Malaysia (Borneo).
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  • Ecuadorian Toadheaded Pitviper (Bothrocophias campbelli). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • With a mouthful of dry moss, a Whitehead's Pygmy Squirrel (Exilisciurus whiteheadi) pauses cautiously to make sure nobody is watching before disappearing into its nest in a rock crevice. Endemic to the island of Borneo, this tiny squirrel lives only in cool montane rainforests.
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  • Madagascan Nightjar (Caprimulgus madagascariensis). Tomasina, Madagascar.
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  • While pythons undoubtedly hold the crown for the largest and most famous constricting snakes of Indonesia, a single genus of boas also occurs in the rainforests of the eastern archipelago. Often overlooked, the Pacific Keel-scaled Boa (Candoia carinata) seldom exceeds a meter in length. It is an ambush predator of frogs and lizards, and like other boas, bears live young rather than laying eggs. Digul River, Papua, Indonesia (New Guinea).
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  • The Twin-barred Flying Snake (Chrysopelea pelias) is the smallest and rarest of Southeast Asia's flying snake species. Like other Chrysopelea, this snake hunts lizards and is almost completely arboreal. Sarawak, Malaysia (Borneo).
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  • Distinguished by its green/yellow eyes, Hose's Bush Frog (Philautus hosii) is endemic to Borneo and considered Near Threatened due to severe habitat loss. Adults are usually found perched on vegetation over streams in lowland rainforest. Sarawak, Malaysia (Borneo).
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  • The dainty White-eared Tree Frog (Feihyla kajau) is endemic to the rainforests of Borneo. Sarawak, Malaysia.
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  • Borneo Anglehead Lizard (Gonocephalus bornensis). Sarawak, Malaysia (Borneo).
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  • The Long-tongued Nectar Bat (Macroglossus minimus) is an important pollinator of many rainforest trees. Here it is feeding on the nectar of banana flowers. Sarawak, Malaysia (Borneo).
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  • Extremely agile on the wing, the Bat Hawk (Macheiramphus alcinus) is a specialist predator of bats and swiftlets, often waiting near cave entrances for its prey to emerge. Sarawak, Malaysia (Borneo).
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  • The Oriental Vine Snake (Ahaetulla prasina) is an agile climber, feeding on small birds and lizards with its excellent binocular vision. Sabah, Malaysia (Borneo).
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  • Mountain Treeshrew (Tupaia montana) feeding at Nepenthes rajah.  Recent research has shown the the world's largest pitcher plant Nepenthes rajah is not exclusively carnivorous. Like the related N. lowii, this species attracts treeshrews by secreting nectar on the undersurface of the lid. These animals frequently leave their droppings in the pitcher, which serves as a valuable nitrogen source in their impoverished mountain habitat. Mount Kinabalu National Park, Sabah, Malaysia (Borneo).
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  • Goodman's Mouse Lemur (Microcebus lehilahytsara), feeding on the nectar of night-blooming Dombeya flowers. Andasibe, Madagascar.
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  • Tree Frog (Litoria rubella). Wasur National Park, Papua, Indonesia (New Guinea).
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  • A young Parson's Chameleon (Calumma parsonii) in the rainforest of Masoala National Park. Antsiranana, Madagascar.
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  • Sri Lanka Giant Squirrel (Ratufa macroura dandolena). Central Province, Sri Lanka.
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  • Endemic to northern Borneo, the Black-and-crimson Pitta (Pitta ussheri) is a shy forest bird which feeds on worms and other terrestrial invertebrates. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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