Chien C. Lee

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  • Plain-pouched Hornbill (Rhyticeros subruficollis), male at nest. Huai Kha Khaeng Wildlife Sanctuary, Thailand.
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  • Crested Serpent Eagle (Spilornis cheela). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • With his crop full of food and water, a male Plain-pouched Hornbill (Rhyticeros subruficollis) returns to his nest in the cavity of a tree wherein the female waits sealed inside. During the nesting period, which can last several months, his mate will be completely dependent on his regular visits to provide her with everything she needs while she incubates the young and raises the chicks. Unfortunately, his duties on this day have been complicated by the annoying attentions of a Greater Racket-tailed Drongo (Dicrurus paradiseus) which has perhaps also been nesting nearby. Although much smaller than the hornbill, drongos are known for their bold and aggressive behavior, especially towards other birds that could be potential predators of their nests. Huai Kha Khaeng Wildlife Sanctuary, Thailand.
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  • Brown-tailed Racer (Gonyosoma oxycephalum). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • The Smooth Otter (Lutrogale perspicillata) is the largest otter species in Southeast Asia, reaching up to 1.3 meters in length. Smooth Otters mate for life and hunt cooperatively for fish. Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Estuarine Crocodile (Crocodylus porosus). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Estuarine Crocodile (Crocodylus porosus). Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Red-bearded Bee-eater (Nyctyornis amictus), male. Tabin Wildlife Reserve, Sabah, Malaysia (Borneo).
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  • The Smooth Otter (Lutrogale perspicillata) is the largest otter species in Southeast Asia, reaching up to 1.3 meters in length. Smooth Otters mate for life and hunt cooperatively for fish. Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • In the coastal forests of northern Borneo, a young Silvery Lutung (Trachypithecus cristatus) is groomed by its mother. Like many leaf monkey species, these highly social primates display brightly colored fur when born that gradually fades after several months into their adult coloration. The exact reason for this adaption is uncertain, but it may help in protection: members of the group can easily distinguish their vulnerable young at a glance, or locate them quickly in the dense foliage of the rainforest canopy. Another interesting theory is that certain predators such as Clouded Leopards, are red/green colorblind, so an orangish pelt may actually be an effective camouflage. Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Borneo Pygmy Elephant (Elephas maximus borneensis). Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Sunda Clouded Leopard (Neofelis diardi), female. Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Bronzeback (Dendrelaphis terrificus). Gorontalo, Sulawesi, Indonesia.
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  • The shy Maleo (Macrocephalon maleo) is a large megapode bird endemic to Sulawesi. They do not build nests but instead bury their eggs in huge pits dug in hot volcanic sand. The precocious young are able to fly and fend for themselves immediately after hatching. Central Sulawesi, Indonesia.
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  • Smith's Litter Frog (Leptobrachium smithi). Krabi, Thailand.
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  • The largest terrestrial and river predator in the world, the Estuarine Crocodile (Crocodylus porosus) can reach lengths of over 6 m and consumes a wide range of animal prey. They are commonly found in mangroves, lagoons, and large rivers along the coastline of Borneo.  Kinabatangan River, Sabah, Malaysia (Borneo).
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  • At only 12cm in length, the Sulawesi Dwarf Kingfisher (Ceyx fallax) is one of the smallest kingfishers. It is a bird of the forest, feeding on small lizards and insects, and is endemic to Sulawesi and a few offshore islands, although scarce and patchily distributed. Gorontalo, Indonesia.
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  • The rare and enigmatic Babirusa (Babyrousa celebensis) occurs only on Sulawesi and a few neighboring islands. Adult males have four conspicuous tusks which are formed from their elongated canine teeth. Gorontalo, Sulawesi, Indonesia.
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  • The Asian Eyebrow-ridge Toad (Duttaphrynus melanostictus) is a common species widespread in South and Southeast Asia.  It has a distinctive large parotoid glands behind the head, which are known to secrete a toxic substance when the toad is disturbed.
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  • Sulawesi Babirusa (Babyrousa celebensis). Gorontalo, Sulawesi.
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  • Closeup portrait of a Tokay Gecko (Gekko gecko). Uthai Thani, Thailand.
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  • Endemic to Sulawesi and the Sula Islands, the Sulawesi Black Pigeon (Turacoena manadensis) inhabits coastal, lowland, and hill forests. Gorontalo, Indonesia.
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  • Terciopelo (Bothrops asper). Cartago, Costa Rica.
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  • Smith's Green-eyed Gecko (Gekko smithii). Kinabatangan River, Sabah, Malaysia (Borneo).
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  • Brown Bullfrog (Kaloula baleata). Sabah, Malaysia (Borneo).
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  • Harlequin Tree Frog (Rhacophorus pardalis). Sabah, Malaysia (Borneo).
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  • A large group of Proboscis Monkeys (Nasalis larvatus) prepares for sleep in a tree overhanging a river. Sabah, Malaysia (Borneo).
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  • The rare and elusive Gurney's Pitta (Pitta gurneyi) is regarded as one of the world's most endangered birds.  Until recently this species was known from less than 20 individuals all occurring in a single reserve in southern Thailand.  Following the discovery of additional populations in southern Myanmar in 2003, the status Gurney's Pitta was downgraded from Critically Endangered to Endangered.  This species occurs only in flat lowland rainforest, a rapidly disappearing habitat in this region due to logging and oil-palm plantations. The adult male bird pictured here sports a leg band from a recent population survey. Krabi, Thailand.
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  • A male Black-naped Monarchs (Hypothymis azurea) at his nest. In these flycatchers both male and female will share equal care in tending to the nestlings. Krabi, Thailand.
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  • An adult Common Butterfly Lizard (Leiolepis belliana) keeps a watchful eye out for predators whilst her young remain at the entrance of the nest burrow.  Nine species of butterfly lizards are known from the drier sandy regions of Southeast Asia, but their life-history is poorly documented. Uthai Thani, Thailand.
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  • Maleo (Macrocephalon maleo), pair digging in black sand for egg laying. Central Sulawesi, Indonesia.
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  • A crystal-clear stream flows through the forest at Khao Nor Chuchi Wildlife Sanctuary.  These spring-fed waters which emerge from limestone rock are highly alkaline and contain high amounts of dissolved calcium carbonate. Krabi, Thailand.
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  • Mixed Deciduous Forest during the beginning of the rainy season in Huai Kha Khaeng Wildlife Sanctuary.
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  • Dense bamboo forest along edges of Huai Kha Khaeng River, consisting mostly of huge stands of Giant Thorny Bamboo (Bambusa bambos). Huai Kha Khaeng Wildlife Sanctuary, Thailand.
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  • Proboscis Monkey (Nasalis larvatus). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • A large male Bornean Clouded Leopard (Neofelis diardi) is caught on camera trap as he patrols his territory in the rainforest of Danum Valley, Sabah. Borneo's largest cat species, the clouded leopard preys on a variety of smaller wildlife ranging from deer and wild boar to monkeys. Sabah, Malaysia (Borneo).
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  • Although sometimes weighing over 9 kg, Indri (Indri indri) can make impressive leaps from tree to tree with their powerful long hind legs. Largest of the living lemurs, this critically-endangered species is an icon for the conservation of Madagascar's beleaguered wildlife, and its wailing song never fails to enchant visitors to their forest realm. Occuring only in the tropical rainforests on the island's eastern side, the Indri's natural range has dwindled due to widespread deforestation and hunting, despite their traditionally sacred status. Tomasina, Madagascar.
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  • Without seeing it move, you could almost mistake the Golden Mantella (Mantella aurantiaca) for a bright orange plastic toy. This is one of Madagascar’s most endangered amphibians and is an icon for conservation of the island’s threatened wildlife. Efforts targeted at protecting this frog’s habitat, coupled with ex-situ breeding programs and reintroduction have helped to protect it from extinction in the wild, but it remains highly threatened and is still known to exist at only two small isolated patches of rainforest. Andasibe, Madagascar.
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  • Without seeing it move, you could almost mistake the Golden Mantella (Mantella aurantiaca) for a bright orange plastic toy. This is one of Madagascar’s most endangered amphibians and is an icon for conservation of the island’s threatened wildlife. Efforts targeted at protecting this frog’s habitat, coupled with ex-situ breeding programs and reintroduction have helped to protect it from extinction in the wild, but it remains highly threatened and is still known to exist at only two small isolated patches of rainforest. Andasibe, Madagascar.
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  • Ground beetle (Chlaenius sabahensis). Sungai Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Lichen Huntsman Spider (Heteropoda boiei) with parasitic biting midges (Ceratopogonidae). Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Sumatran Two-tailed Spider (Hersilia sumatrana). Kinabatangan Wildlife Sanctuary, Sabah, Malaysia (Borneo).
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  • Some of the most noticeable spiders of the Southeast Asian tropical rainforests are the huntsmans (family Sparassidae); not only do they reach impressive sizes and are often colorful but because they don’t use webs for their prey capture they can easily be seen hunting in understory vegetation at night. Some, such as this Lichen Huntsman (Pandercetes sp.) are harder to find than most due to their cryptic camouflage. This female specimen rests atop her equally camouflaged egg sac which has been neatly woven into the fold of a dead leaf. Tabin Wildlife Reserve, Sabah, Malaysia (Borneo).
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  • Bornean Orangutan (Pongo pygmaeus), infant. Sarawak, Malaysia (Borneo).
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  • Although usually considered a terrestrial species, the Striped Kukri Snake (Oligodon octolineatus) is an excellent climber and can occasionally be seen climbing tree trunks. Sarawak, Malaysia (Borneo).
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  • A female Bornean Orang-utan (Pongo pygmaeus) with her young infant. Semengoh Forest Reserve, Sarawak, Malaysia (Borneo).
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  • Borneo Horned Frog (Pelobatrachus nasutus), female, with Matang Narrow-mouthed Frog (Microhyla nepenthicola) - comparison between one of the largest and smallest frogs in Borneo. Sarawak, Malaysia (Borneo).
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  • Red-crowned Barbet (Psilopogon rafflesii), a frugivorous bird from the rainforests of Southeast Asia. Sarawak, Malaysia (Borneo).
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  • Although usually considered a terrestrial species, the Striped Kukri Snake (Oligodon octolineatus) is an excellent climber and can occasionally be seen climbing tree trunks. Sarawak, Malaysia (Borneo).
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  • Greater Sac-winged Bat (Saccopteryx bilineata). Colón, Panama.
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  • White-spotted Cat Snake (Boiga drapiezii), green morph. Sarawak, Malaysia (Borneo).
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  • Hidden in the dense leaf litter of the rainforest floor, an Amazonian Horned Frog (Ceratophrys cornuta) lies in wait for its next meal. With a mouth wider than the length of its body, and a voracious appetite to boot, these frogs can consume prey as large as small reptiles and rodents. This species is widely distributed in the Amazon Basin, but nowhere particularly abundant. Yasuní National Park, Ecuador.
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  • Bornean Opposite-fingered Tree Frog (Feihyla inexpectata), first photographic record for this species in Sarawak. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Spectacled Flying Fox (Pteropus conspicillatus). Waigeo Island, West Papua, Indonesia.
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  • Sun Glass Frog (Hyalinobatrachium aureoguttatum), male hiding on the underside of a leaf. The pattern on the back of this species mimics a cluster of the frog's eggs, which help to deter attacks by predatory wasps. Esmeraldas, Ecuador.
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  • Larut Bush Frog (Philautus larutensis), male. Mulu National Park, Sarawak, Malaysia (Borneo).
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  • Ivohimanita Madagascar Frog (Mantidactylus majori), male guarding eggs. Ranomafana National Park, Madagascar.
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  • Few creatures possess a mastery of camouflage the way Madagascar’s leaf-tailed geckos do. This Mossy Leaf-tailed Gecko (Uroplatus sikorae) sleeps by day on a rainforest liana, its outline obscured by frilled dermal flaps that line its body. By always resting in a downward-facing position they may be able to quickly catch unsuspecting insects that climb up from the ground and also conceal their reflective lidless eyes from predators lurking above. Ranomafana National Park, Madagascar.
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  • Smallest of all the anteaters, the Silky Anteater (Cyclopes didactylus) is seldom seen because it spends much of its life high in the rainforest canopy, often curled up as an indistinguishable ball of fluff. They have no teeth and can only defend themselves by means of their razor sharp sickle-like fore-claws, which are usually used for tearing open ant nests. Females bear a single youngster at a time, which is carried on their back until large enough to feed on its own. Esmeraldas, Ecuador.
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  • Sri Lankan Grey Langur (Semnopithecus priam thersites). Yala National Park, Sri Lanka.
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  • Matang Narrow-mouthed Frog (Microhyla nepenthicola), in amplexus.  This is the smallest frog in the Old World and it rears its young exclusively in the fluid of pitcher plants. Sarawak, Malaysia (Borneo).
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  • Endemic to the mountains of northern Borneo, the Kinabalu Horned Frog (Pelobatrachus baluensis) breeds only in clear rocky streams with cold water. By day they lie camouflaged among the leaf litter on the forest floor, but they emerge at night to call for mates. Sabah, Malaysia (Borneo).
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  • Nycticebus menagensis. One of four species of Slow Loris found in Borneo. All Slow Loris are nocturnal arboreal primates which feed on small animals, insects and soft fruits. Sarawak, Malaysia (Borneo).
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  • Ecuadorian Toadheaded Pitviper (Bothrocophias campbelli). Mashpi Amagusa Reserve, Pichincha, Ecuador.
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  • The Twin-barred Flying Snake (Chrysopelea pelias) is the smallest and rarest of Southeast Asia's flying snake species. Like other Chrysopelea, this snake hunts lizards and is almost completely arboreal. Sarawak, Malaysia (Borneo).
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  • Malayan Flat-shelled Turtle (Notochelys platynota). Sarawak, Malaysia (Borneo).
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  • Sabah Bamboo Pitviper (Trimeresurus sabahi). Sarawak, Malaysia (Borneo).
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  • The Long-tongued Nectar Bat (Macroglossus minimus) is an important pollinator of many rainforest trees. Here it is feeding on the nectar of banana flowers. Sarawak, Malaysia (Borneo).
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  • Without a single sighting in nearly 90 years, the Borneo Rainbow Toad (Ansonia latidisca) was considered possibly extinct and listed by Conservation International as one of the "10 Most Wanted Amphibians". The species was rediscovered in 2011 by a team of herpetologists from the University of Malaysia Sarawak, although it remains extremely rare and little is known of its ecology or behavior. This brings a glimmer of hope in a time of global mass amphibian extinctions. Sarawak, Malaysia (Borneo).
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  • Flat-nosed Pitviper (Craspedocephalus puniceus). West Java, Indonesia.
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  • The Black-bearded Tomb Bat (Taphozous melanopogon) roosts in caves and large rock crevices, emerging at night to feed on insects high above the forest canopy. It is found throughout much of mainland Southeast Asia as well as Borneo, Sumatra, and Java. It's echolocation calls while flying are audible as high pitched clicks. Sarawak, Malaysia (Borneo).
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  • Extremely agile on the wing, the Bat Hawk (Macheiramphus alcinus) is a specialist predator of bats and swiftlets, often waiting near cave entrances for its prey to emerge. Sarawak, Malaysia (Borneo).
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  • Tree Frog (Litoria rubella). Wasur National Park, South Papua, Indonesia (New Guinea).
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  • When threatened, the Knob-tailed Gecko (Nephrurus levis) raises its body off the ground to appear larger in size. Cape Range National Park, Western Australia.
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  • A male Whitehead's Broadbill (Calyptomena whiteheadi) returns to its nest where it passes food to the hungry chick. Both male and female care for the nestlings, providing insects and fruit, as well as removing fecal pellets. This scarcely-seen species is the largest of all the green broadbills (reaching 27cm in length) and is endemic to the montane forests of Borneo. Sabah, Malaysia.
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  • Endemic to northern Borneo, the Black-and-crimson Pitta (Pitta ussheri) is a shy forest bird which feeds on worms and other terrestrial invertebrates. Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Threatened by habitat loss and hunting pressure, the Crested Black Macaque (Macaca nigra nigra) is now listed as Critically Endangered. This species lives only in forested areas at the northernmost tip of the island of Sulawesi. North Sulawesi, Indonesia.
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  • Endemic to the mountains of central Sulawesi, the stream-breeding Loka Flying Frog (Rhacophorus monticola) is highly variable in coloration. South Sulawesi, Indonesia.
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  • This Collared Nightjar (Gactornis enarratus), a Madagascar endemic, hides itself among the leaf litter of the forest floor while it sleeps. Placed in its own genus, this species is distinct from all other nightjars (family Caprimulgidae) by its plumage, eggs, and behavior. Mantadia National Park, Madagascar.
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  • Hikida's Forest Dragon (Hypsilurus hikidanus). Central Papua, Indonesia (New Guinea).
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  • Usambara Eyelash Viper (Atheris ceratophora). Udzungwa Mountains, Tanzania.
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  • Sunda Frogmouth (Batrachostomus cornutus), female. Sabah, Malaysia (Borneo).
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  • Banded Malaysian Coralsnake (Calliophis intestinalis). Sarawak, Malaysia (Borneo).
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  • Polynesian Rat (Rattus exulans). Sarawak, Malaysia (Borneo).
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  • Boophis luciae, amplexing pair. Ranomafana National Park, Madagascar.
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  • Black-headed Cat Snake (Boiga nigriceps). Danum Valley Conservation Area, Sabah, Malaysia (Borneo).
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  • Sarayacu Tree Frog (Dendropsophus parviceps). Orellana, Ecuador.
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  • The Rufous Piculet (Sasia abnormis) is the smallest woodpecker in Borneo.  Here a bird is making its territorial call by drumming on a spiny rattan petiole. Sabah, Malaysia (Borneo).
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  • The Palawan Peacock Pheasant (Polyplectron napoleonis) is a shy ground-dwelling bird endemic to the rainforests the island of Palawan, Philippines.
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  • Spotted Cuscus (Spilocuscus maculatus chrysorrhous), juvenile. Central Papua, Indonesia (New Guinea).
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  • Three-banded Bridled Snake (Lycodon tristrigatus). Sarawak, Malaysia (Borneo).
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  • Pitta-like Ground-roller (Atelornis pittoides) with earthworm. Ranomafana National Park, Madagascar.
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  • Guatemalan Yellow Grosbeak (Pheucticus chrysopeplus aurantiacus), vocalising. El Triunfo Biosphere Reserve, Chiapas, Mexico.
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  • Long-tailed Sylph (Aglaiocercus kingii). Papallacta, Ecuador.
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  • The mimicry of a dead leaf is so precise for the Satanic Leaf-tailed Gecko (Uroplatus phantasticus), they can often hide effectively in plain sight. It has been untested to what degree these geckos actively select a substrate that matches their individual patterns, but U. phantasticus appears to typically hide among dead foliage which suits it perfectly. Ranomafana National Park, Madagascar.
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  • Owston's Palm Civet (Chrotogale owstoni), a nocturnal carnivore from Laos and Vietnam, is a highly vulnerable species due to illegal poaching. Captive. Endangered Primate Rescue Centre, Cuc Phuong National Park, Vietnam.
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  • Madagascar Plover (Charadrius thoracicus). Toliara, Madagascar.
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  • By means of its extensive skin membranes, the Sunda Colugo (Galeopterus variegatus borneanus), is capable of gliding impressive distances between trees. This adult is carrying a young offspring on its underside. Bearing no relation to true lemurs, colugos are in their own unique order and are only distantly related to primates and treeshrews. They are nocturnal, feeding on leaves, flowers, and sap. Recent research on morphological and genetic variations indicate that the Bornean and Javan subspecies may merit recognition as distinct species. Sarawak, Malaysia (Borneo).
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  • Lesser Bare-backed fruit Bat (Dobsonia minor), feeding on fig. Nimbokrang, Papua, Indonesia (New Guinea).
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  • Marañón White-fronted Capuchin (Cebus yuracus). Yasuní National Park, Ecuador.
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